Agoseris heterophylla |
Agoseris heterophylla var. cryptopleura |
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annual agoseris, annual false dandelion, mountain dandelion |
California agoseris, mountain dandelion |
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Habit | Annuals. | |||||||||
Stems | 0 or 1 (erect, 0–5 cm). |
0 or 1 (internodes to 1/2 times lengths of subtending leaves). |
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Leaves | mostly erect, sometimes prostrate; petioles not purplish, margins glabrous or ciliate; blades usually oblanceolate to spatulate, rarely linear, 1–25 cm, margins entire or lobed; lobes 2–3 pairs, linear to spatulate, spreading to antrorse, lobules mostly 0, glabrous or densely hairy. |
erect to spreading; blades (2–)5–15(–24) cm × (1.5–)2–10(–16) mm, mostly toothed to lobed, rarely entire, teeth or lobes (2–)3–5 pairs, faces usually densely, ± uniformly hairy, rarely glabrous. |
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Peduncles | elongating after flowering, 3–60 cm in fruit, glabrous or glabrate, or basally puberulent and apically hairy to tomentose, sometimes stipitate-glandular. |
6–60 cm, lengths mostly 1.5–4 times leaves at flowering, 2–5 times length of leaves in fruit, glabrous or glabrate, or apically hairy to tomentose. |
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Involucres | cylindric to hemispheric, 1–2 cm in fruit. |
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Receptacles | epaleate. |
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Florets | 5–100(–300); corollas yellow, tubes 1–5 mm, ligules 2–15 × 1–3 mm; anthers 1–4 mm. |
20–100(–300); corollas much surpassing phyllaries at flowering, tubes 2–5 mm, ligules 10–15 × 2–3 mm; anthers 2–4 mm. |
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Phyllaries | in 2–3 series, green or medially rosy purple, sometimes purple-black spotted or tipped, subequal to unequal, margins glabrous or ciliate, faces usually puberulent to villous, mostly stipitate-glandular, sometimes glabrous; outer erect or spreading, adaxially usually villous to lanate, sometimes glabrous; inner erect, ± elongating after flowering. |
green or medially rosy purple, sometimes spotted, margins glabrous or ciliate, faces mostly hairy, occasionally glabrous or villous, stipitate-glandular, with translucent, yellowish or purple-septate, often purple-tipped hairs or eglandular with whitish-opaque hairs; outer erect to spreading, adaxially glabrous or pubescent and eglandular. |
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Heads | (7–)10–18 mm wide (flowering). |
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Cypselae | 7–16 mm, bodies mostly fusiform to obconic, sometimes tumid, 2–5(–10) mm, beaks 5–11 mm, lengths 1–4 times bodies, ribs 0 or alate, straight to strongly undulate, uniform or diminishing proximally; pappus bristles in 2–3 series, 4–9 mm. |
outermost strongly differing from inner, highly variable as to color, shape, ornamentation, and pubescence, ribs flattened to alate, or 0, straight to strongly undulate, often strongly diminishing proximally. |
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2n | = 18. |
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Agoseris heterophylla |
Agoseris heterophylla var. cryptopleura |
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Phenology | Flowering Mar–Sep. | |||||||||
Habitat | Mesic to dry habitats in grasslands, chaparral, oak woodlands, and open pine forests | |||||||||
Elevation | 150–2100 m (500–6900 ft) | |||||||||
Distribution |
AZ; CA; ID; MT; NM; NV; OR; UT; WA; BC; nw Mexico (including Guadalupe Island) [Introduced in Europe (Sweden)]
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CA |
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Discussion | Varieties 3 (3 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Variety cryptopleura occurs on hills and ranges surrounding the Great Central Valley (it appears to be absent from the valley itself). It is ± sympatric with var. heterophylla, except that var. cryptopleura almost completely supplants var. heterophylla in the south Coast Ranges of California. Older floras treated this taxon as var. californica; more recent works have not recognized it at all. K. L. Chambers (1963b) demonstrated that var. cryptopleura is strictly allogamous and is not self-fertile. Varieties heterophylla and cryptopleura are differentiated almost entirely by corolla size, which appears to be correlated with breeding system; otherwise the two exhibit almost complete morphologic overlap. Corolla size in depauperate specimens of var. cryptopleura approaches that found in robust specimens of var. heterophylla. The two varieties still separate on corollas longer than the phyllaries in var. cryptopleura versus subequaling the phyllaries in var. heterophylla. The degree of introgression, if any, between var. cryptopleura and var. heterophylla is not known. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 19, p. 332. | FNA vol. 19, p. 333. | ||||||||
Parent taxa | Asteraceae > tribe Cichorieae > Agoseris | Asteraceae > tribe Cichorieae > Agoseris > Agoseris heterophylla | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Macrorhynchus heterophyllus | A. californica, A. heterophylla subsp. californica, A. heterophylla var. californica, A. heterophylla var. crenulata, A. heterophylla var. turgida, A. major, Cryptopleura californica | ||||||||
Name authority | (Nuttall) Greene: Pittonia 2: 178. (1891) | Greene: Pittonia 2: 179. (1891) | ||||||||
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