Agalinis neoscotica |
Agalinis oligophylla |
|
---|---|---|
Middleton false foxglove, Nova scotia agalinis, Nova scotia false foxglove |
ridgestem false foxglove |
|
Stems | simple or branched, 5–30(–47) cm; branches spreading-ascending, obtusely quadrangular proximally to quadrangular with wings on angles distally, glabrous or sparsely scabridulous. |
simple or branched, 30–90(–110) cm; branches spreading-ascending, quadrangular, with siliceous ridges on angles, scabridulous. |
Leaves | spreading; blade linear-elliptic to elliptic, 6–40 x 0.8–5 mm, not fleshy, margins entire, abaxial midvein sparsely scabrous, adaxial surface scabrous; axillary fascicles absent or shorter than subtending leaves. |
ascending; blade subulate, elliptic, or filiform, (1–)4–13 x 0.3–1.1 mm, margins entire, heavily siliceous, scabridulous, adaxial surface siliceous, scabridulous; axillary fascicles absent. |
Inflorescences | racemes, flowers 2 per node; bracts longer than pedicels. |
racemiform-paniculate, with short multinoded floriferous branches, some with pseudoterminal flowers, flowers 1 or 2 per node; bracts shorter than or sometimes equal to pedicels. |
Pedicels | spreading-ascending, (0.8–)1.5–5.3(–6.5) mm, glabrous. |
ascending, 4–16 mm, +/- scabridulous. |
Flowers | calyx hemispheric-campanulate, tube 1.5–3 mm, glabrous, lobes triangular-lanceolate, 1.1–4(–5) mm, unequal, mid adaxial shortest; corolla pale to dark pink, with 2 yellow lines and red spots in abaxial throat, or lines faint or absent, (8–)10–15 mm, throat pilose externally and villous within across bases and sinus of adaxial lobes, lobes: abaxial slightly spreading to projected, adaxial slightly spreading or projected distal to corolla mouth, 1.6–6 mm, unequal, pilose or densely so externally; proximal and distal anthers parallel to filaments, pollen sacs 1–2 mm; style included, (3–)4–7.5 mm. |
calyx campanulate to funnelform, tube 2.5–3.2 mm, glabrous, lobes subulate to triangular-subulate, 0.1–1 mm; corolla pink to dark pink, with 2 yellow lines and red spots in abaxial throat, 15–25 mm, throat pilose externally and villous within across bases and sinus of adaxial lobes, lobes: abaxial spreading, adaxial reflexed-spreading, 4–7(–9) mm, glabrous externally; proximal anthers parallel to filaments, distal oblique or perpendicular to filaments, pollen sacs 1.8–2.6 mm; style exserted, 6–11 mm. |
Capsules | globular, 5–6 mm. |
globular, 4–6 mm. |
Seeds | brown, 0.9–1.5 mm. |
tan to yellow, 0.5–0.9 mm. |
2n | = 28. |
= 26. |
Agalinis neoscotica |
Agalinis oligophylla |
|
Phenology | Flowering late Jul–early Sep. | Flowering late Sep–mid Nov. |
Habitat | Sandy or peaty soils, margins of pools, lakes, bogs, estuaries, and marshes, sand flats, dune hollows, ditches. | Moist to dry longleaf pine savannas, edges of pine plantations, dry roadsides, chalky outcrops, seepage slopes of clay roadsides. |
Elevation | 0–10 m. (0–0 ft.) | 0–300 m. (0–1000 ft.) |
Distribution |
ME; NB; NS
|
AL; AR; LA; MS; TN; TX |
Discussion | Agalinis neoscotica is rare in Maine, occurring only in Washington County on shores and tidal pools, and in adjacent eastern New Brunswick; it is most abundant in southwestern Nova Scotia and is reported from neighboring Grand Manan Island and Sable Island on the extreme northeastern edge of the range of Agalinis. Features that characterize Agalinis neoscotica are narrow tubular corolla throats with red spots in two rows on the two yellow lines (the lines sometimes absent); corolla lobes that either project forward or are only slightly spreading; relatively small anthers, all held parallel to the filaments; two-lobed stigmas, obvious in live plants but rarely on dried specimens; both anthers and stigma included within the corolla throat; calyx in which the middle adaxial lobe is shortest and curved toward the corolla; bracts longer than both the pedicels and flowers they subtend and, sometimes, longer than the main stem leaves; and leaves with narrowed bases that terminate at a creaselike abscission zone. Agalinis neoscotica has a mixed mating system (H. M. Stewart et al. 1996). Self-pollination occurs in bud but delayed pollen germination allows for out-crossing. Stewart and J. M. Canne (1998) presented data that showed A. neoscotica flower development and morphology differ from those of A. purpurea. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Plants of Agalinis oligophylla are rare in the easternmost portions of the range; they are abundant farther west into Texas. Agalinis oligophylla is often the last species of Agalinis to begin flowering, usually in late September, sometimes not until mid October and often remains in flower well into November. Unbranched plants of A. oligophylla can be confused with A. aphylla, with which it often shares the habitat. Agalinis oligophylla can be distinguished from A. aphylla by the longer leaves, more diffusely branched inflorescence, glabrous style, and the much later flowering period of A. oligophylla. Also, on the older buds of A. oligophylla the corolla is globular, but in A. aphylla the corolla on older buds is obovoid. Corollas of A. oligophylla are asymmetric; the adaxial side of the corolla throat is conspicuously shorter than the upcurved abaxial side of the corolla. The buds, pedicels, and capsules of A. oligophylla turn dark brown or blackish, and corollas turn dark purplish or pinkish brown when dried. These dark structures contrast strikingly with the pale green to gray (or reddish) branches and leaves. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 17, p. 548. | FNA vol. 17, p. 549. |
Parent taxa | Orobanchaceae > Agalinis | Orobanchaceae > Agalinis |
Sibling taxa | ||
Synonyms | Gerardia neoscotica, A. paupercula var. neoscotica, A. purpurea var. neoscotica | Gerardia aphylla var. grandiflora, G. microphylla |
Name authority | (Greene) Fernald: Rhodora 23: 139. (1921) | Pennell: Bull. Torrey Bot. Club 40: 432. (1913) |
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