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strigose lotus

red and yellow pea, scrub deervetch or lotus, Wright's deervetch

Habit Herbs, annual, usually mat-forming, sometimes cespitose (ascending and bushy), green to grayish, 0.3–5 dm, not or ± fleshy, strigillose, hirsute, canescent-tomentose, or scantily pubescent; taprooted. Herbs, perennial, cespitose, erect or sprawling, greenish or grayish, 2–4(–8) dm, not fleshy, usually strigose to hirsute, rarely canescent; from woody caudices.
Stems

1–20+, procumbent or decumbent to ascending, branched basally, herbaceous, slender, leafy.

1–20+, procumbent to ascending, branched proximally, stiff, leafy, base without persistent leaves.

Leaves

irregularly pinnate;

stipules glandlike;

subsessile;

rachis 3–20 mm, flattened;

leaflets 4–10, blades unequal, obovate to oblanceolate to linear-oblong or oblong, apex acute to obtuse or truncate, surfaces ± densely tomentose to glabrate.

palmate, often ± dimorphic (proximal with broader leaflets, distal with filiform ones), sometimes subtending axillary clusters of filiform leaflets;

stipules ovate;

proximal short-petiolate, medial and distal subsessile or sessile;

rachis absent;

leaflets 3–6, blades obovate (proximal) or oblanceolate to linear (distal), apex usually acute, sometimes obtuse, surfaces ± densely villous to strigose.

Inflorescences

1–3-flowered.

1 or 2-flowered.

Peduncles

ascending or reflexed, upturned, 3–25 mm, shorter to longer than leaves (often elongated in fruit);

bract absent or 1–3-foliolate, usually subtending umbel.

ascending to deflexed, 0–30 mm, usually longer, sometimes shorter, than leaves;

bract absent or (when pedunculate) unifoliolate, distal.

Flowers

(5–)6–10(–12) mm;

calyx 3–5.5 mm, tube ± sparsely strigillose or glabrous, lobes subulate;

corolla yellow (sometimes banner orangish abaxially), turning orange or reddish, claws ± equaling calyx tube, banner implicate-ascending or remaining closely implicate, wings longer than keel;

style corneously thickened, marked by color, straight, puberulent or glabrous.

10–15(–18) mm;

calyx 5–7.5 mm, tube villous to strigose, lobes subulate to setaceous;

corolla yellow with back of banner red, turning reddish, claws shorter than calyx tube, banner implicate-ascending, wings longer than keel;

style straight, glabrous.

Legumes

persistent, exserted, erect or divergent, brown, straight to ± curved distally, compressed, not or slightly constricted, incompletely septate, linear-oblong, 10–35 × 2–3 mm, thinly leathery, apex initially short-beaked, dehiscent, smooth, margins smooth, thin, glabrous or ± strigose.

persistent, exserted, divergent or declined, reddish or grayish brown, straight, turgid, slightly constricted, incompletely septate, linear-oblong, (17–)20–35 × 1.5–3(–3.5) mm, leathery, apex short-beaked, dehiscent, smooth, margins smooth, thickened, sparsely strigose to glabrate.

Seeds

5–10, greenish to brown, ± mottled, ovoid to cuboid or globose, rugulose or granular.

4–7, greenish to dark reddish brown, ± mottled, oblong, smooth.

2n

= 14.

Acmispon strigosus

Acmispon wrightii

Phenology Flowering late spring–summer.
Habitat Canyon slopes, mesas, washes, ± dry hillsides, sandy-loam, sandy, or gravelly soils, sometimes saline, pine-oak-Douglas-fir, pine or aspen forests, oak-pinyon-Cercocarpus or pinyon-juniper woodlands, juniper-oak grasslands, desert grasslands or scrub, riparian woodlands, roadsides.
Elevation (1500–)1700–3200 m. ((4900–)5600–10500 ft.)
Distribution
sw United States; n Mexico
[BONAP county map]
from FNA
AZ; CO; NM; NV; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

D. Isely (1981) discussed the variation in Acmispon strigosus (as Lotus), distinguishing three varieties, two rather distinct but with intergrading phases, and one apparently more transitional, although distinct enough to be recognized. These varieties were only noted for California in recent treatments (for example, L. Brouillet 2012, following D. Isely 1993). Nonetheless, the typical (for example, excluding intergrading phases) varieties represent recognizable morphologies that also have geographic distinction. Specimens representing intergradation seem to occur throughout the range of overlap of these varieties, sometimes rendering determination difficult.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Acmispon wrightii is reported to hybridize with A. mearnsii var. mearnsii, A. rigidus, and A. utahensis (A. M. Ottley 1944; D. Isely 1981), as well as A. plebeius (Isely).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Herbs evidently hirsute, especially at apex, greenish.
var. hirtellus
1. Herbs usually strigillose or canescent-tomentose to scantly pubescent, rarely strigillose, green or cinereous.
→ 2
2. Stems ascending or decumbent, not markedly succulent, usually appearing green, sometimes cinereous; leaflet blades linear-oblong, apices acute to obtuse; corolla opening; California.
var. strigosus
2. Stems prostrate, succulent, cinereous(–greenish); leaflet blades oblong, apices obtuse or truncate; corolla not opening; Arizona, California, Nevada.
var. tomentellus
Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Acmispon Fabaceae > subfam. Faboideae > Acmispon
Sibling taxa
A. americanus, A. argophyllus, A. argyraeus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. denticulatus, A. glaber, A. grandiflorus, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. tomentosus, A. utahensis, A. wrangelianus, A. wrightii
A. americanus, A. argophyllus, A. argyraeus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. denticulatus, A. glaber, A. grandiflorus, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. strigosus, A. tomentosus, A. utahensis, A. wrangelianus
Subordinate taxa
A. strigosus var. hirtellus, A. strigosus var. strigosus, A. strigosus var. tomentellus
Synonyms Hosackia strigosa, Anisolotus strigosus, Lotus strigosus, Ottleya strigosa Hosackia wrightii, Anisolotus wrightii, Lotus wrightii, Ottleya wrightii
Name authority (Nuttall) Brouillet: J. Bot. Res. Inst. Texas 2: 392. (2008) (A. Gray) Brouillet: J. Bot. Res. Inst. Texas 2: 392. (2008)
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