Acmispon strigosus |
Acmispon wrangelianus |
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strigose lotus |
California lotus, Chilean bird's-foot trefoil, Chilean trefoil, hairy lotus |
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Habit | Herbs, annual, usually mat-forming, sometimes cespitose (ascending and bushy), green to grayish, 0.3–5 dm, not or ± fleshy, strigillose, hirsute, canescent-tomentose, or scantily pubescent; taprooted. | Herbs, annual, cespitose, grayish green, 0.5–3 dm, ± not fleshy, sparsely villous to glabrate; taprooted. | ||||||||
Stems | 1–20+, procumbent or decumbent to ascending, branched basally, herbaceous, slender, leafy. |
1–20+, usually procumbent (when small, unbranched, erect or ascending), diffusely branched proximally, herbaceous, leafy. |
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Leaves | irregularly pinnate; stipules glandlike; subsessile; rachis 3–20 mm, flattened; leaflets 4–10, blades unequal, obovate to oblanceolate to linear-oblong or oblong, apex acute to obtuse or truncate, surfaces ± densely tomentose to glabrate. |
irregularly pinnate; stipules glandlike or absent; petiolate; rachis 5–15 mm, flattened; leaflets (3 or)4(or 5), usually 2 on one side and 2 terminal, blades elliptic to obovate (lateral sometimes asymmetric), apex usually obtuse, sometimes acute, surfaces sparsely long-ciliate, pubescent. |
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Inflorescences | 1–3-flowered. |
1-flowered. |
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Peduncles | ascending or reflexed, upturned, 3–25 mm, shorter to longer than leaves (often elongated in fruit); bract absent or 1–3-foliolate, usually subtending umbel. |
ascending, 0–2 mm, shorter than leaves; bract absent. |
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Flowers | (5–)6–10(–12) mm; calyx 3–5.5 mm, tube ± sparsely strigillose or glabrous, lobes subulate; corolla yellow (sometimes banner orangish abaxially), turning orange or reddish, claws ± equaling calyx tube, banner implicate-ascending or remaining closely implicate, wings longer than keel; style corneously thickened, marked by color, straight, puberulent or glabrous. |
5–9 mm; calyx 2.5–5 mm, tube pubescent to villous, lobes lanceolate; corolla yellow, turning red, claws shorter than calyx tube, banner implicate-ascending, with irregular or inrolled margins, wings shorter than keel; style ± curved, glabrous. |
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Legumes | persistent, exserted, erect or divergent, brown, straight to ± curved distally, compressed, not or slightly constricted, incompletely septate, linear-oblong, 10–35 × 2–3 mm, thinly leathery, apex initially short-beaked, dehiscent, smooth, margins smooth, thin, glabrous or ± strigose. |
persistent, exserted, erect or spreading, tawny to brown, straight, compressed, slightly constricted, ± septate, oblong, 10–18 × 2.2–3 mm, thinly leathery, apex abruptly downward angled and curved, dehiscent, smooth, margins smooth, thin, pubescent (to glabrate). |
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Seeds | 5–10, greenish to brown, ± mottled, ovoid to cuboid or globose, rugulose or granular. |
3–7, olive green to brown, not mottled, asymmetric-reniform, smooth. |
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2n | = 12. |
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Acmispon strigosus |
Acmispon wrangelianus |
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Phenology | Flowering (late winter–)spring(–early summer). | |||||||||
Habitat | Coastal bluffs, hills, open or rocky slopes, chaparral, coastal scrub, oak woodlands or savannas, grasslands, stream banks, vernal ponds, gravelly, sandy, or clayey soils, bare areas, burnt or disturbed areas, roadsides. | |||||||||
Elevation | 0–1200(–2200) m. (0–3900(–7200) ft.) | |||||||||
Distribution | sw United States; n Mexico |
CA; OR
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Discussion | Varieties 3 (3 in the flora). D. Isely (1981) discussed the variation in Acmispon strigosus (as Lotus), distinguishing three varieties, two rather distinct but with intergrading phases, and one apparently more transitional, although distinct enough to be recognized. These varieties were only noted for California in recent treatments (for example, L. Brouillet 2012, following D. Isely 1993). Nonetheless, the typical (for example, excluding intergrading phases) varieties represent recognizable morphologies that also have geographic distinction. Specimens representing intergradation seem to occur throughout the range of overlap of these varieties, sometimes rendering determination difficult. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Acmispon wrangelianus is widespread in California, except in the Mojave Desert, barely entering Oregon in Jackson County; it is possibly introduced in the Modoc Plateau and Sonoran Desert. It was found in 1911 on ballast in Multnomah County, Oregon, but does not appear to have become established there. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 11. | FNA vol. 11. | ||||||||
Parent taxa | Fabaceae > subfam. Faboideae > Acmispon | Fabaceae > subfam. Faboideae > Acmispon | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Hosackia strigosa, Anisolotus strigosus, Lotus strigosus, Ottleya strigosa | Lotus wrangelianus, Anisolotus wrangeliana, Hosackia wrangeliana, L. subpinnatus var. wrangelianus | ||||||||
Name authority | (Nuttall) Brouillet: J. Bot. Res. Inst. Texas 2: 392. (2008) | (Fisher & C. A. Meyer) D. D. Sokoloff: Taxon 48: 58. (1999) | ||||||||
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