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strigose lotus

Utah bird's-foot trefoil, Utah deervetch or lotus

Habit Herbs, annual, usually mat-forming, sometimes cespitose (ascending and bushy), green to grayish, 0.3–5 dm, not or ± fleshy, strigillose, hirsute, canescent-tomentose, or scantily pubescent; taprooted. Herbs, perennial, cespitose, greenish or grayish (base), 1–5 dm, not fleshy, puberulent to sparsely strigose or glabrate; rhizomatous caudex from a taproot.
Stems

1–20+, procumbent or decumbent to ascending, branched basally, herbaceous, slender, leafy.

1–10+, decumbent to ascending or erect, branched sparsely, stiff, leafy, base without persistent leaves.

Leaves

irregularly pinnate;

stipules glandlike;

subsessile;

rachis 3–20 mm, flattened;

leaflets 4–10, blades unequal, obovate to oblanceolate to linear-oblong or oblong, apex acute to obtuse or truncate, surfaces ± densely tomentose to glabrate.

palmate, homomorphic;

stipules ovate;

sessile;

rachis absent;

leaflets (2–)4–6, blades linear-oblanceolate to oblanceolate, apex acute to obtuse, surfaces villous to strigose.

Inflorescences

1–3-flowered.

(1 or)2–5(or 6)-flowered.

Peduncles

ascending or reflexed, upturned, 3–25 mm, shorter to longer than leaves (often elongated in fruit);

bract absent or 1–3-foliolate, usually subtending umbel.

ascending to deflexed, curved to erect, (10–)20–50(–80) mm, longer than leaves;

bract 1–3-foliolate, subtending umbel.

Flowers

(5–)6–10(–12) mm;

calyx 3–5.5 mm, tube ± sparsely strigillose or glabrous, lobes subulate;

corolla yellow (sometimes banner orangish abaxially), turning orange or reddish, claws ± equaling calyx tube, banner implicate-ascending or remaining closely implicate, wings longer than keel;

style corneously thickened, marked by color, straight, puberulent or glabrous.

8–15 mm;

calyx 5–7.5 mm, tube villous, lobes subulate;

corolla yellow with red-backed banner, often suffused with red, turning orange, claws shorter than calyx tube, banner recurved ca. 45°, pandurate, wings longer than keel (nearly equaling banner);

style straight, glabrous.

Legumes

persistent, exserted, erect or divergent, brown, straight to ± curved distally, compressed, not or slightly constricted, incompletely septate, linear-oblong, 10–35 × 2–3 mm, thinly leathery, apex initially short-beaked, dehiscent, smooth, margins smooth, thin, glabrous or ± strigose.

persistent, exserted, spreading or deflexed, reddish to grayish brown, ± straight, turgid, not constricted, incompletely septate, linear-oblong, (14–)25–35 × 1.5–3.5 mm, leathery, apex short-beaked, dehiscent, smooth, lustrous, margins keeled, strigillose or glabrous.

Seeds

5–10, greenish to brown, ± mottled, ovoid to cuboid or globose, rugulose or granular.

(1–)3–6, olive green to dark brown, mottled, oblong, smooth.

Acmispon strigosus

Acmispon utahensis

Phenology Flowering spring(–summer).
Habitat Open places, dry, stony or sandy soils, sagebrush, desert scrub, pinyon-juniper woodlands, chaparral, oak woodlands, yellow pine or spruce-aspen forests, riparian communities.
Elevation (1500–)1700–2900 m. ((4900–)5600–9500 ft.)
Distribution
sw United States; n Mexico
[BONAP county map]
from FNA
AZ; NV; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

D. Isely (1981) discussed the variation in Acmispon strigosus (as Lotus), distinguishing three varieties, two rather distinct but with intergrading phases, and one apparently more transitional, although distinct enough to be recognized. These varieties were only noted for California in recent treatments (for example, L. Brouillet 2012, following D. Isely 1993). Nonetheless, the typical (for example, excluding intergrading phases) varieties represent recognizable morphologies that also have geographic distinction. Specimens representing intergradation seem to occur throughout the range of overlap of these varieties, sometimes rendering determination difficult.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Acmispon utahensis is nearly restricted to Utah, barely penetrating into Arizona and Nevada. It is encountered on the plateaus of the intermountain region. It is reported to hybridize with A. rigidus and A. wrightii (A. M. Ottley 1944).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Herbs evidently hirsute, especially at apex, greenish.
var. hirtellus
1. Herbs usually strigillose or canescent-tomentose to scantly pubescent, rarely strigillose, green or cinereous.
→ 2
2. Stems ascending or decumbent, not markedly succulent, usually appearing green, sometimes cinereous; leaflet blades linear-oblong, apices acute to obtuse; corolla opening; California.
var. strigosus
2. Stems prostrate, succulent, cinereous(–greenish); leaflet blades oblong, apices obtuse or truncate; corolla not opening; Arizona, California, Nevada.
var. tomentellus
Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Acmispon Fabaceae > subfam. Faboideae > Acmispon
Sibling taxa
A. americanus, A. argophyllus, A. argyraeus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. denticulatus, A. glaber, A. grandiflorus, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. tomentosus, A. utahensis, A. wrangelianus, A. wrightii
A. americanus, A. argophyllus, A. argyraeus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. denticulatus, A. glaber, A. grandiflorus, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. strigosus, A. tomentosus, A. wrangelianus, A. wrightii
Subordinate taxa
A. strigosus var. hirtellus, A. strigosus var. strigosus, A. strigosus var. tomentellus
Synonyms Hosackia strigosa, Anisolotus strigosus, Lotus strigosus, Ottleya strigosa Lotus utahensis, Ottleya utahensis
Name authority (Nuttall) Brouillet: J. Bot. Res. Inst. Texas 2: 392. (2008) (Ottley) Brouillet: J. Bot. Res. Inst. Texas 2: 392. (2008)
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