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strigose lotus

Habit Herbs, annual, usually mat-forming, sometimes cespitose (ascending and bushy), green to grayish, 0.3–5 dm, not or ± fleshy, strigillose, hirsute, canescent-tomentose, or scantily pubescent; taprooted. Herbs, perennial, mat-forming, greenish, 0.5–3 dm, not fleshy, densely hirsute; from woody caudices.
Stems

1–20+, procumbent or decumbent to ascending, branched basally, herbaceous, slender, leafy.

1–20+, procumbent or decumbent, branched, herbaceous, leafy, proximally covered by small, persistent leaves.

Leaves

irregularly pinnate;

stipules glandlike;

subsessile;

rachis 3–20 mm, flattened;

leaflets 4–10, blades unequal, obovate to oblanceolate to linear-oblong or oblong, apex acute to obtuse or truncate, surfaces ± densely tomentose to glabrate.

pinnate to subpalmate (distal);

stipules conic;

proximal short-petiolate or sessile, distal subsessile to short-petiolate;

rachis 0–4 mm, sometimes flattened;

leaflets (3 or)4 or 5(or 6), blades obovate to oblanceolate (proximal ± orbiculate), apex acute to rounded, surfaces villous to tomentose.

Inflorescences

1–3-flowered.

1–3(–5)-flowered.

Peduncles

ascending or reflexed, upturned, 3–25 mm, shorter to longer than leaves (often elongated in fruit);

bract absent or 1–3-foliolate, usually subtending umbel.

ascending or reflexed, upturned, (5–)20–50 mm, longer than leaves;

bract absent (reduced to gland) or 1(–3)-foliolate, distal.

Flowers

(5–)6–10(–12) mm;

calyx 3–5.5 mm, tube ± sparsely strigillose or glabrous, lobes subulate;

corolla yellow (sometimes banner orangish abaxially), turning orange or reddish, claws ± equaling calyx tube, banner implicate-ascending or remaining closely implicate, wings longer than keel;

style corneously thickened, marked by color, straight, puberulent or glabrous.

(10–)13–22 mm;

calyx 4.3–7 mm, tube densely villous, lobes subulate;

corolla cream to orange-yellow, rose-tinted, claws shorter than calyx tube, banner obliquely ascending, wings longer than banner and keel, auriculate, spurred;

style ± straight, glabrous.

Legumes

persistent, exserted, erect or divergent, brown, straight to ± curved distally, compressed, not or slightly constricted, incompletely septate, linear-oblong, 10–35 × 2–3 mm, thinly leathery, apex initially short-beaked, dehiscent, smooth, margins smooth, thin, glabrous or ± strigose.

persistent, exserted, erect, reddish brown, straight, turgid, not constricted, imperfectly septate, linear-oblong, 15–30 × (2–)2.5–3 mm, ± leathery, apex short-beaked, dehiscent, smooth, margins smooth, strigillose.

Seeds

5–10, greenish to brown, ± mottled, ovoid to cuboid or globose, rugulose or granular.

5–8, olive green to brown, ± mottled, oblong, smooth.

2n

= 14.

Acmispon strigosus

Acmispon neomexicanus

Phenology Flowering (late winter–)spring.
Habitat Washes, canyons, stream banks, sandy or clayey soils, dry, gravelly and rocky slopes, riparian woodlands, desert or mesquite grasslands, desert slope scrub, grassy mountain foothills, chaparral, pinyon-oak-juniper woodlands, oak savannas.
Elevation 1200–2600 m. (3900–8500 ft.)
Distribution
sw United States; n Mexico
[BONAP county map]
from FNA
AZ; NM; Mexico (Chihuahua, Durango, Sonora)
Discussion

Varieties 3 (3 in the flora).

D. Isely (1981) discussed the variation in Acmispon strigosus (as Lotus), distinguishing three varieties, two rather distinct but with intergrading phases, and one apparently more transitional, although distinct enough to be recognized. These varieties were only noted for California in recent treatments (for example, L. Brouillet 2012, following D. Isely 1993). Nonetheless, the typical (for example, excluding intergrading phases) varieties represent recognizable morphologies that also have geographic distinction. Specimens representing intergradation seem to occur throughout the range of overlap of these varieties, sometimes rendering determination difficult.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Acmispon neomexicanus occurs in southeastern Arizona and adjacent southwestern New Mexico. It is reported to hybridize with A. plebeius and A. rigidus (A. M. Ottley 1944; D. Isely 1981). Anisolotus greenei Wooton & Standley and Ottleya greenei (Wooton & Standley) D. D. Sokoloff are superfluous illegitimate names; Hosackia mollis Greene (1885, not Nuttall 1838), and Lotus mollis Greene (1890, not Balfour f. 1882) are illegitimate names that pertain here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Herbs evidently hirsute, especially at apex, greenish.
var. hirtellus
1. Herbs usually strigillose or canescent-tomentose to scantly pubescent, rarely strigillose, green or cinereous.
→ 2
2. Stems ascending or decumbent, not markedly succulent, usually appearing green, sometimes cinereous; leaflet blades linear-oblong, apices acute to obtuse; corolla opening; California.
var. strigosus
2. Stems prostrate, succulent, cinereous(–greenish); leaflet blades oblong, apices obtuse or truncate; corolla not opening; Arizona, California, Nevada.
var. tomentellus
Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Acmispon Fabaceae > subfam. Faboideae > Acmispon
Sibling taxa
A. americanus, A. argophyllus, A. argyraeus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. denticulatus, A. glaber, A. grandiflorus, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. tomentosus, A. utahensis, A. wrangelianus, A. wrightii
A. americanus, A. argophyllus, A. argyraeus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. denticulatus, A. glaber, A. grandiflorus, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. strigosus, A. tomentosus, A. utahensis, A. wrangelianus, A. wrightii
Subordinate taxa
A. strigosus var. hirtellus, A. strigosus var. strigosus, A. strigosus var. tomentellus
Synonyms Hosackia strigosa, Anisolotus strigosus, Lotus strigosus, Ottleya strigosa Lotus neomexicanus, A. greenei, Anisolotus mollis, A. neomexicanus, Hosackia greenei, H. neomexicana, Lotus greenei, Ottleya mollis
Name authority (Nuttall) Brouillet: J. Bot. Res. Inst. Texas 2: 392. (2008) (Greene) Brouillet: Phytoneuron 2020-29: 2. (2020)
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