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strigose lotus

coastal bird's-foot trefoil, coastal lotus

Habit Herbs, annual, usually mat-forming, sometimes cespitose (ascending and bushy), green to grayish, 0.3–5 dm, not or ± fleshy, strigillose, hirsute, canescent-tomentose, or scantily pubescent; taprooted. Herbs, annual, cespitose, green, 0.5–3.5(–5) dm, ± fleshy, glabrous or strigillose; taprooted.
Stems

1–20+, procumbent or decumbent to ascending, branched basally, herbaceous, slender, leafy.

1–20, procumbent to ascending, branched basally, herbaceous, leafy.

Leaves

irregularly pinnate;

stipules glandlike;

subsessile;

rachis 3–20 mm, flattened;

leaflets 4–10, blades unequal, obovate to oblanceolate to linear-oblong or oblong, apex acute to obtuse or truncate, surfaces ± densely tomentose to glabrate.

irregularly pinnate;

stipules glandlike;

sessile or subsessile to short-petiolate;

rachis 8–20(–35) mm, flattened;

leaflets (3–)5–7, blades unequal, obovate to ± orbiculate, apex obtuse, surfaces glabrous or ± strigillose.

Inflorescences

1–3-flowered.

1–5-flowered.

Peduncles

ascending or reflexed, upturned, 3–25 mm, shorter to longer than leaves (often elongated in fruit);

bract absent or 1–3-foliolate, usually subtending umbel.

erect then spreading or declined, 3–15 mm, shorter to longer than leaves, sometimes branched, slender;

bract absent or 1–3-foliolate, subtending umbel.

Flowers

(5–)6–10(–12) mm;

calyx 3–5.5 mm, tube ± sparsely strigillose or glabrous, lobes subulate;

corolla yellow (sometimes banner orangish abaxially), turning orange or reddish, claws ± equaling calyx tube, banner implicate-ascending or remaining closely implicate, wings longer than keel;

style corneously thickened, marked by color, straight, puberulent or glabrous.

2.5–8(–11) mm;

calyx 1.2–4.5 mm, tube strigose to villous, lobes lanceolate;

corolla bright yellow or orange-yellow, claws shorter than to equaling calyx tube, banner implicate-ascending, wings symmetric, equaling or shorter than keel;

style curved, glabrous.

Legumes

persistent, exserted, erect or divergent, brown, straight to ± curved distally, compressed, not or slightly constricted, incompletely septate, linear-oblong, 10–35 × 2–3 mm, thinly leathery, apex initially short-beaked, dehiscent, smooth, margins smooth, thin, glabrous or ± strigose.

persistent, exserted, ascending or spreading, tawny to brown, straight or curved, ± compressed, scarcely or distinctly constricted, not septate, narrowly oblong, 10–30 × 3–4 mm, leathery, apex short hook-beaked, dehiscent, margins smooth, thin, glabrous or sparsely strigillose.

Seeds

5–10, greenish to brown, ± mottled, ovoid to cuboid or globose, rugulose or granular.

5–9, brown to dark olive green, not mottled, oblong-ovoid, smooth.

Acmispon strigosus

Acmispon maritimus

Distribution
sw United States; n Mexico
[BONAP county map]
w United States; nw Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

D. Isely (1981) discussed the variation in Acmispon strigosus (as Lotus), distinguishing three varieties, two rather distinct but with intergrading phases, and one apparently more transitional, although distinct enough to be recognized. These varieties were only noted for California in recent treatments (for example, L. Brouillet 2012, following D. Isely 1993). Nonetheless, the typical (for example, excluding intergrading phases) varieties represent recognizable morphologies that also have geographic distinction. Specimens representing intergradation seem to occur throughout the range of overlap of these varieties, sometimes rendering determination difficult.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Herbs evidently hirsute, especially at apex, greenish.
var. hirtellus
1. Herbs usually strigillose or canescent-tomentose to scantly pubescent, rarely strigillose, green or cinereous.
→ 2
2. Stems ascending or decumbent, not markedly succulent, usually appearing green, sometimes cinereous; leaflet blades linear-oblong, apices acute to obtuse; corolla opening; California.
var. strigosus
2. Stems prostrate, succulent, cinereous(–greenish); leaflet blades oblong, apices obtuse or truncate; corolla not opening; Arizona, California, Nevada.
var. tomentellus
1. Flowers 6–8(–11) mm; calyces 4–4.5 mm, lobes 1.5–2.2(–2.5) mm; legumes (15–)20–30 mm, scarcely constricted between seeds.
var. maritimus
1. Flowers 2.5–4 mm; calyces 1.2–2 mm, lobes 0.2–1 mm; legumes 10–15 mm, distinctly constricted between seeds.
var. brevivexillus
Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Acmispon Fabaceae > subfam. Faboideae > Acmispon
Sibling taxa
A. americanus, A. argophyllus, A. argyraeus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. denticulatus, A. glaber, A. grandiflorus, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. tomentosus, A. utahensis, A. wrangelianus, A. wrightii
A. americanus, A. argophyllus, A. argyraeus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. denticulatus, A. glaber, A. grandiflorus, A. haydonii, A. intricatus, A. junceus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. strigosus, A. tomentosus, A. utahensis, A. wrangelianus, A. wrightii
Subordinate taxa
A. strigosus var. hirtellus, A. strigosus var. strigosus, A. strigosus var. tomentellus
A. maritimus var. brevivexillus, A. maritimus var. maritimus
Synonyms Hosackia strigosa, Anisolotus strigosus, Lotus strigosus, Ottleya strigosa Hosackia maritima, Anisolotus maritimus, Lotus salsuginosus
Name authority (Nuttall) Brouillet: J. Bot. Res. Inst. Texas 2: 392. (2008) (Nuttall) D. D. Sokoloff: Ann. Bot. Fenn. 37: 129. (2000)
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