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strigose lotus

rush deervetch, rush deervetch or trefoil, rush lotus

Habit Herbs, annual, usually mat-forming, sometimes cespitose (ascending and bushy), green to grayish, 0.3–5 dm, not or ± fleshy, strigillose, hirsute, canescent-tomentose, or scantily pubescent; taprooted. Subshrubs (often flowering 1st year, appearing annual), bushy or wiry and prostrate, sometimes robust, brownish, 0.8–4 dm, not fleshy, strigillose to glabrate; from woody caudices.
Stems

1–20+, procumbent or decumbent to ascending, branched basally, herbaceous, slender, leafy.

1–20+, prostrate to ascending, branched, ± woody, wiry or stout, often remotely leafy.

Leaves

irregularly pinnate;

stipules glandlike;

subsessile;

rachis 3–20 mm, flattened;

leaflets 4–10, blades unequal, obovate to oblanceolate to linear-oblong or oblong, apex acute to obtuse or truncate, surfaces ± densely tomentose to glabrate.

irregularly subpalmate;

stipules glandlike;

sessile or subsessile to short-petiolate;

rachis 0–9 mm, sometimes flattened;

leaflets 3–5, blades obovate to oblanceolate, apex obtuse to acute, or mucronulate, surfaces ± strigillose.

Inflorescences

1–3-flowered.

(1 or)2–8-flowered.

Peduncles

ascending or reflexed, upturned, 3–25 mm, shorter to longer than leaves (often elongated in fruit);

bract absent or 1–3-foliolate, usually subtending umbel.

ascending, slender, sometimes secondarily branched, 1–25 mm, usually shorter, sometimes longer, than leaves;

bract absent or (when pedunculate) unifoliolate, distal.

Flowers

(5–)6–10(–12) mm;

calyx 3–5.5 mm, tube ± sparsely strigillose or glabrous, lobes subulate;

corolla yellow (sometimes banner orangish abaxially), turning orange or reddish, claws ± equaling calyx tube, banner implicate-ascending or remaining closely implicate, wings longer than keel;

style corneously thickened, marked by color, straight, puberulent or glabrous.

6–8 mm;

calyx 3–5 mm, tube strigillose, lobes triangular to deltate;

corolla yellow, tinged red, fading orange, claws longer than calyx tube, banner implicate-ascending, wings shorter than keel (and other petals);

style gradually or abruptly upcurved, glabrous.

Legumes

persistent, exserted, erect or divergent, brown, straight to ± curved distally, compressed, not or slightly constricted, incompletely septate, linear-oblong, 10–35 × 2–3 mm, thinly leathery, apex initially short-beaked, dehiscent, smooth, margins smooth, thin, glabrous or ± strigose.

persistent, moderately to well exserted, ascending to divergent, tawny to brown, strongly arched, curved to 90°, sometimes nearly straight, turgid, not constricted, not septate, linear-oblong, 6–8 × 1–2 mm, leathery, apex beak recurved 80–360° or irregularly contorted, nearly as long as body, indehiscent, transversely ridged, margins keeled, rugose, strigillose or glabrous.

Seeds

5–10, greenish to brown, ± mottled, ovoid to cuboid or globose, rugulose or granular.

1 or 2, olive to reddish brown, mottled, elongate, curved, smooth.

Acmispon strigosus

Acmispon junceus

Distribution
sw United States; n Mexico
[BONAP county map]
California
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

D. Isely (1981) discussed the variation in Acmispon strigosus (as Lotus), distinguishing three varieties, two rather distinct but with intergrading phases, and one apparently more transitional, although distinct enough to be recognized. These varieties were only noted for California in recent treatments (for example, L. Brouillet 2012, following D. Isely 1993). Nonetheless, the typical (for example, excluding intergrading phases) varieties represent recognizable morphologies that also have geographic distinction. Specimens representing intergradation seem to occur throughout the range of overlap of these varieties, sometimes rendering determination difficult.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Herbs evidently hirsute, especially at apex, greenish.
var. hirtellus
1. Herbs usually strigillose or canescent-tomentose to scantly pubescent, rarely strigillose, green or cinereous.
→ 2
2. Stems ascending or decumbent, not markedly succulent, usually appearing green, sometimes cinereous; leaflet blades linear-oblong, apices acute to obtuse; corolla opening; California.
var. strigosus
2. Stems prostrate, succulent, cinereous(–greenish); leaflet blades oblong, apices obtuse or truncate; corolla not opening; Arizona, California, Nevada.
var. tomentellus
1. Stems usually stout, prostrate to ascending; peduncles 1–5 mm; legumes moderately exserted.
var. junceus
1. Stems often wiry, usually prostrate; peduncles (3–)8–25 mm; legumes well exserted.
var. biolettii
Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Acmispon Fabaceae > subfam. Faboideae > Acmispon
Sibling taxa
A. americanus, A. argophyllus, A. argyraeus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. denticulatus, A. glaber, A. grandiflorus, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. tomentosus, A. utahensis, A. wrangelianus, A. wrightii
A. americanus, A. argophyllus, A. argyraeus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. denticulatus, A. glaber, A. grandiflorus, A. haydonii, A. intricatus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. strigosus, A. tomentosus, A. utahensis, A. wrangelianus, A. wrightii
Subordinate taxa
A. strigosus var. hirtellus, A. strigosus var. strigosus, A. strigosus var. tomentellus
A. junceus var. biolettii, A. junceus var. junceus
Synonyms Hosackia strigosa, Anisolotus strigosus, Lotus strigosus, Ottleya strigosa Hosackia juncea, Lotus junceus, Syrmatium junceum
Name authority (Nuttall) Brouillet: J. Bot. Res. Inst. Texas 2: 392. (2008) (Bentham) Brouillet: J. Bot. Res. Inst. Texas 2: 391. (2008)
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