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strigose lotus

California broom, common deerweed, deer weed, deervetch, western bird's-foot trefoil

Habit Herbs, annual, usually mat-forming, sometimes cespitose (ascending and bushy), green to grayish, 0.3–5 dm, not or ± fleshy, strigillose, hirsute, canescent-tomentose, or scantily pubescent; taprooted. Subshrubs, often bushy, sometimes mat-forming, robust, green, 5–20 dm, not fleshy, glabrous or finely strigose; from woody caudices.
Stems

1–20+, procumbent or decumbent to ascending, branched basally, herbaceous, slender, leafy.

1–30+, usually erect to spreading, rarely procumbent, branched basally, herbaceous, slender, striate, sometimes ± woody, remotely leafy, usually deciduous mid-season, foliage developing after rain.

Leaves

irregularly pinnate;

stipules glandlike;

subsessile;

rachis 3–20 mm, flattened;

leaflets 4–10, blades unequal, obovate to oblanceolate to linear-oblong or oblong, apex acute to obtuse or truncate, surfaces ± densely tomentose to glabrate.

± pinnate (distally 3-foliolate);

stipules glandlike;

subsessile to petiolate;

rachis 2–8(–10) mm, not flattened;

leaflets 3–6(or 7), blades elliptic to lanceolate, apex acute, surfaces strigillose to glabrate.

Inflorescences

1–3-flowered.

(1 or)2–7-flowered, in axil of distal leaves, spaced or congested.

Peduncles

ascending or reflexed, upturned, 3–25 mm, shorter to longer than leaves (often elongated in fruit);

bract absent or 1–3-foliolate, usually subtending umbel.

ascending, 0–2 mm, shorter than leaves;

bract absent.

Flowers

(5–)6–10(–12) mm;

calyx 3–5.5 mm, tube ± sparsely strigillose or glabrous, lobes subulate;

corolla yellow (sometimes banner orangish abaxially), turning orange or reddish, claws ± equaling calyx tube, banner implicate-ascending or remaining closely implicate, wings longer than keel;

style corneously thickened, marked by color, straight, puberulent or glabrous.

7–12 mm;

calyx 2.5–5 mm, tube usually glabrous, sometimes sparsely strigillose, lobes short-subulate;

corolla yellow, turning orange or red, claws slightly longer than calyx tube, banner implicate-ascending to 90°, wings shorter than or equaling keel;

style curved, glabrous.

Legumes

persistent, exserted, erect or divergent, brown, straight to ± curved distally, compressed, not or slightly constricted, incompletely septate, linear-oblong, 10–35 × 2–3 mm, thinly leathery, apex initially short-beaked, dehiscent, smooth, margins smooth, thin, glabrous or ± strigose.

persistent, much exserted, divergent or pendent, greenish to reddish brown, curved to ± straight, turgid, slightly constricted, not septate, linear-oblong, 10–15 × 1–2 mm, leathery, apex long-beaked, indehiscent, smooth, margins thickened, smooth, strigillose or glabrate.

Seeds

5–10, greenish to brown, ± mottled, ovoid to cuboid or globose, rugulose or granular.

(1 or)2, olive to brown, not mottled, elongate-oblong, smooth.

Acmispon strigosus

Acmispon glaber

Distribution
sw United States; n Mexico
[BONAP county map]
nw Mexico; California
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

D. Isely (1981) discussed the variation in Acmispon strigosus (as Lotus), distinguishing three varieties, two rather distinct but with intergrading phases, and one apparently more transitional, although distinct enough to be recognized. These varieties were only noted for California in recent treatments (for example, L. Brouillet 2012, following D. Isely 1993). Nonetheless, the typical (for example, excluding intergrading phases) varieties represent recognizable morphologies that also have geographic distinction. Specimens representing intergradation seem to occur throughout the range of overlap of these varieties, sometimes rendering determination difficult.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

D. Isely (1981) reported a potential hybrid between Acmispon glaber (variety not specified) and A. argophyllus (variety not specified). Lotus glaber (Vogel) Greene 1890 (not Miller 1768) is an illegitimate name that pertains here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Herbs evidently hirsute, especially at apex, greenish.
var. hirtellus
1. Herbs usually strigillose or canescent-tomentose to scantly pubescent, rarely strigillose, green or cinereous.
→ 2
2. Stems ascending or decumbent, not markedly succulent, usually appearing green, sometimes cinereous; leaflet blades linear-oblong, apices acute to obtuse; corolla opening; California.
var. strigosus
2. Stems prostrate, succulent, cinereous(–greenish); leaflet blades oblong, apices obtuse or truncate; corolla not opening; Arizona, California, Nevada.
var. tomentellus
1. Flowers 7–12 mm; keel ± equaling wings.
var. glaber
1. Flowers 8–9(–10) mm; keel usually longer than wings.
var. brevialatus
Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Acmispon Fabaceae > subfam. Faboideae > Acmispon
Sibling taxa
A. americanus, A. argophyllus, A. argyraeus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. denticulatus, A. glaber, A. grandiflorus, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. tomentosus, A. utahensis, A. wrangelianus, A. wrightii
A. americanus, A. argophyllus, A. argyraeus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. denticulatus, A. grandiflorus, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. strigosus, A. tomentosus, A. utahensis, A. wrangelianus, A. wrightii
Subordinate taxa
A. strigosus var. hirtellus, A. strigosus var. strigosus, A. strigosus var. tomentellus
A. glaber var. brevialatus, A. glaber var. glaber
Synonyms Hosackia strigosa, Anisolotus strigosus, Lotus strigosus, Ottleya strigosa Syrmatium glabrum, Anisolotus glaber, Hosackia glabra
Name authority (Nuttall) Brouillet: J. Bot. Res. Inst. Texas 2: 392. (2008) (Vogel) Brouillet: J. Bot. Res. Inst. Texas 2: 389. (2008) — (as glabrus)
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