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strigose lotus

pine deervetch or lotus

Habit Herbs, annual, usually mat-forming, sometimes cespitose (ascending and bushy), green to grayish, 0.3–5 dm, not or ± fleshy, strigillose, hirsute, canescent-tomentose, or scantily pubescent; taprooted. Herbs, perennial (seldom flowering first year), mat-forming or cespitose, diffuse, low, greenish gray (young growth cinereous), 0.5–1 dm (3–12 dm wide), not fleshy, ±villous to glabrate, hairs curved; woody-taprooted.
Stems

1–20+, procumbent or decumbent to ascending, branched basally, herbaceous, slender, leafy.

1–20+, procumbent, branched, wiry, slender, leafy.

Leaves

irregularly pinnate;

stipules glandlike;

subsessile;

rachis 3–20 mm, flattened;

leaflets 4–10, blades unequal, obovate to oblanceolate to linear-oblong or oblong, apex acute to obtuse or truncate, surfaces ± densely tomentose to glabrate.

irregularly pinnate to subpalmate;

stipules glandlike;

subsessile to petiolate;

rachis (1–)2–10(–15) mm, not or ± flattened;

leaflets 3–5, blades obovate to elliptic, apex acute to obtuse, surfaces villous, hairs curved.

Inflorescences

1–3-flowered.

(1–)3–12-flowered.

Peduncles

ascending or reflexed, upturned, 3–25 mm, shorter to longer than leaves (often elongated in fruit);

bract absent or 1–3-foliolate, usually subtending umbel.

ascending, 3–15(–30 proximally) mm, shorter than leaves;

bract 1–3-foliolate, subtending umbel.

Flowers

(5–)6–10(–12) mm;

calyx 3–5.5 mm, tube ± sparsely strigillose or glabrous, lobes subulate;

corolla yellow (sometimes banner orangish abaxially), turning orange or reddish, claws ± equaling calyx tube, banner implicate-ascending or remaining closely implicate, wings longer than keel;

style corneously thickened, marked by color, straight, puberulent or glabrous.

4.5–8(–10) mm;

calyx (3.5–)4–7 mm, tube sparsely villosulous, lobes subulate, subequal;

corolla yellow, reddening, usually drying dark, claws shorter than calyx tube, banner implicate-ascending, wings longer than keel;

style curved, glabrous.

Legumes

persistent, exserted, erect or divergent, brown, straight to ± curved distally, compressed, not or slightly constricted, incompletely septate, linear-oblong, 10–35 × 2–3 mm, thinly leathery, apex initially short-beaked, dehiscent, smooth, margins smooth, thin, glabrous or ± strigose.

deciduous with calyx, exserted, declined or deflexed, tawny, strongly arched, turgid, not constricted, not septate, ovoid-ellipsoid, 6–10 ×1.8–2.2 mm, leathery, apex long, tapering to curved, slender beak, indehiscent, veined, margins keeled, rugose, strigillose.

Seeds

5–10, greenish to brown, ± mottled, ovoid to cuboid or globose, rugulose or granular.

1 or 2 (or 3), brown, not mottled, narrowly oblong-reniform, smooth.

Acmispon strigosus

Acmispon decumbens

Distribution
sw United States; n Mexico
[BONAP county map]
w North America; nw Mexico
Discussion

Varieties 3 (3 in the flora).

D. Isely (1981) discussed the variation in Acmispon strigosus (as Lotus), distinguishing three varieties, two rather distinct but with intergrading phases, and one apparently more transitional, although distinct enough to be recognized. These varieties were only noted for California in recent treatments (for example, L. Brouillet 2012, following D. Isely 1993). Nonetheless, the typical (for example, excluding intergrading phases) varieties represent recognizable morphologies that also have geographic distinction. Specimens representing intergradation seem to occur throughout the range of overlap of these varieties, sometimes rendering determination difficult.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

D. Isely (1981) reported potential hybrids of Acmispon decumbens (variety not specified) with A. argophyllus var. argophyllus and A. glaber var. glaber.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Herbs evidently hirsute, especially at apex, greenish.
var. hirtellus
1. Herbs usually strigillose or canescent-tomentose to scantly pubescent, rarely strigillose, green or cinereous.
→ 2
2. Stems ascending or decumbent, not markedly succulent, usually appearing green, sometimes cinereous; leaflet blades linear-oblong, apices acute to obtuse; corolla opening; California.
var. strigosus
2. Stems prostrate, succulent, cinereous(–greenish); leaflet blades oblong, apices obtuse or truncate; corolla not opening; Arizona, California, Nevada.
var. tomentellus
1. Leaf rachises (3–)5–10(–15) mm; inflorescences (3–)5–12-flowered; flowers 4.5–8(–10) mm; calyces (3.5–)4–7 mm, lobes 0.5–1.5(–3) mm.
var. decumbens
1. Leaf rachises (1–)2–5 mm; inflorescences (1–)3–5(–8)-flowered; flowers 4.5–8(–9) mm; calyces 6–7 mm, lobes 0.5–1.5(–2) mm.
var. davidsonii
Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Acmispon Fabaceae > subfam. Faboideae > Acmispon
Sibling taxa
A. americanus, A. argophyllus, A. argyraeus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. denticulatus, A. glaber, A. grandiflorus, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. tomentosus, A. utahensis, A. wrangelianus, A. wrightii
A. americanus, A. argophyllus, A. argyraeus, A. brachycarpus, A. cytisoides, A. dendroideus, A. denticulatus, A. glaber, A. grandiflorus, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. strigosus, A. tomentosus, A. utahensis, A. wrangelianus, A. wrightii
Subordinate taxa
A. strigosus var. hirtellus, A. strigosus var. strigosus, A. strigosus var. tomentellus
A. decumbens var. davidsonii, A. decumbens var. decumbens
Synonyms Hosackia strigosa, Anisolotus strigosus, Lotus strigosus, Ottleya strigosa Hosackia decumbens, Anisolotus decumbens, Syrmatium decumbens
Name authority (Nuttall) Brouillet: J. Bot. Res. Inst. Texas 2: 392. (2008) (Bentham) Govaerts: Skvortsovia 4(3): 76. (2018)
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