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strigose lotus

Bentham's broom, Bentham's deerweed

Habit Herbs, annual, usually mat-forming, sometimes cespitose (ascending and bushy), green to grayish, 0.3–5 dm, not or ± fleshy, strigillose, hirsute, canescent-tomentose, or scantily pubescent; taprooted. Subshrubs, mat-forming or cespitose, diffusely ascending, low, greenish, 1–8 dm, 50–130 dm diam., not fleshy, glabrous or strigose (young foliage); from woody caudices.
Stems

1–20+, procumbent or decumbent to ascending, branched basally, herbaceous, slender, leafy.

1–20+, procumbent to ± ascending, branched, flexuous-wiry or thick, ± woody, leafy.

Leaves

irregularly pinnate;

stipules glandlike;

subsessile;

rachis 3–20 mm, flattened;

leaflets 4–10, blades unequal, obovate to oblanceolate to linear-oblong or oblong, apex acute to obtuse or truncate, surfaces ± densely tomentose to glabrate.

irregularly pinnate to subpalmate, distal mostly 3-foliolate;

stipules glandlike;

subsessile to short-petiolate;

rachis 1–8 mm, ± flattened;

leaflets 3–5, blades obovate to elliptic, apex obtuse, surfaces glabrous or glabrate.

Inflorescences

1–3-flowered.

3–10-flowered.

Peduncles

ascending or reflexed, upturned, 3–25 mm, shorter to longer than leaves (often elongated in fruit);

bract absent or 1–3-foliolate, usually subtending umbel.

ascending, (1–)2–12(–25) mm, longer than leaves;

bract absent or 1- or 2-foliolate, usually subtending umbel.

Flowers

(5–)6–10(–12) mm;

calyx 3–5.5 mm, tube ± sparsely strigillose or glabrous, lobes subulate;

corolla yellow (sometimes banner orangish abaxially), turning orange or reddish, claws ± equaling calyx tube, banner implicate-ascending or remaining closely implicate, wings longer than keel;

style corneously thickened, marked by color, straight, puberulent or glabrous.

8–10 mm;

calyx 3.5–6 mm, tube sparsely strigillose to glabrate, lobes subulate;

corolla white to pinkish (banner) or yellow (wings cream), banner often red-striate or red-backed, keel reddish or purplish, claws shorter than calyx tube, banner reflexed to 90°, wings slightly longer than keel;

style upcurved, sparsely strigillose.

Legumes

persistent, exserted, erect or divergent, brown, straight to ± curved distally, compressed, not or slightly constricted, incompletely septate, linear-oblong, 10–35 × 2–3 mm, thinly leathery, apex initially short-beaked, dehiscent, smooth, margins smooth, thin, glabrous or ± strigose.

persistent, mostly included, divergent or ascending, brown, arched, turgid, not or slightly constricted, not septate, linear-oblong, 7–10 × 1–2 mm, leathery, apex tapering to exserted, slender, curved or straight beak, indehiscent, smooth, margins thickened, smooth, glabrate, beak strigillose.

Seeds

5–10, greenish to brown, ± mottled, ovoid to cuboid or globose, rugulose or granular.

1 or 2, olive brown, mottled, subcylindric, smooth.

Acmispon strigosus

Acmispon cytisoides

Phenology Flowering spring (early fall).
Habitat Coastal dunes, slopes, and bluffs, cliffs, coastal scrub, chaparral, Monterrey pine forests, exposed slopes, ridges, landslides, gulches, stream banks, sandy or clayey soils, eroded granite, shale, sometimes serpentine outcrops, burnt chaparral, disturbed areas, roadsides.
Elevation 0–1000 m. (0–3300 ft.)
Distribution
sw United States; n Mexico
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

D. Isely (1981) discussed the variation in Acmispon strigosus (as Lotus), distinguishing three varieties, two rather distinct but with intergrading phases, and one apparently more transitional, although distinct enough to be recognized. These varieties were only noted for California in recent treatments (for example, L. Brouillet 2012, following D. Isely 1993). Nonetheless, the typical (for example, excluding intergrading phases) varieties represent recognizable morphologies that also have geographic distinction. Specimens representing intergradation seem to occur throughout the range of overlap of these varieties, sometimes rendering determination difficult.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Acmispon cytisoides occurs in the Central Coast, San Francisco Bay Area, and Outer South Coast Ranges. D. Isely (1981) reported possible hybrids with A. glaber var. glaber (as Lotus scoparius var. perplexans Hoover).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Herbs evidently hirsute, especially at apex, greenish.
var. hirtellus
1. Herbs usually strigillose or canescent-tomentose to scantly pubescent, rarely strigillose, green or cinereous.
→ 2
2. Stems ascending or decumbent, not markedly succulent, usually appearing green, sometimes cinereous; leaflet blades linear-oblong, apices acute to obtuse; corolla opening; California.
var. strigosus
2. Stems prostrate, succulent, cinereous(–greenish); leaflet blades oblong, apices obtuse or truncate; corolla not opening; Arizona, California, Nevada.
var. tomentellus
Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Acmispon Fabaceae > subfam. Faboideae > Acmispon
Sibling taxa
A. americanus, A. argophyllus, A. argyraeus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. denticulatus, A. glaber, A. grandiflorus, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. tomentosus, A. utahensis, A. wrangelianus, A. wrightii
A. americanus, A. argophyllus, A. argyraeus, A. brachycarpus, A. decumbens, A. dendroideus, A. denticulatus, A. glaber, A. grandiflorus, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. strigosus, A. tomentosus, A. utahensis, A. wrangelianus, A. wrightii
Subordinate taxa
A. strigosus var. hirtellus, A. strigosus var. strigosus, A. strigosus var. tomentellus
Synonyms Hosackia strigosa, Anisolotus strigosus, Lotus strigosus, Ottleya strigosa Hosackia cytisoides, H. cytisoides var. rubescens, Lotus benthamii, Syrmatium cytisoides
Name authority (Nuttall) Brouillet: J. Bot. Res. Inst. Texas 2: 392. (2008) (Bentham) Brouillet: J. Bot. Res. Inst. Texas 2: 389. (2008)
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