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strigose lotus

canyon bird's-foot trefoil

Habit Herbs, annual, usually mat-forming, sometimes cespitose (ascending and bushy), green to grayish, 0.3–5 dm, not or ± fleshy, strigillose, hirsute, canescent-tomentose, or scantily pubescent; taprooted. Herbs, perennial, mat-forming, silvery or gray, 1–3 dm, not fleshy, sericeous to canescent; from woody caudices.
Stems

1–20+, procumbent or decumbent to ascending, branched basally, herbaceous, slender, leafy.

1–20+, procumbent or decumbent to diffusely ascending, branched, herbaceous, slender, leafy.

Leaves

irregularly pinnate;

stipules glandlike;

subsessile;

rachis 3–20 mm, flattened;

leaflets 4–10, blades unequal, obovate to oblanceolate to linear-oblong or oblong, apex acute to obtuse or truncate, surfaces ± densely tomentose to glabrate.

palmate or irregularly subpinnate;

stipules glandlike;

subsessile to short-petiolate;

rachis 1–4 mm, ± flattened;

leaflets 3–5, blades obovate to oblanceolate, apex usually obtuse, sometimes acute, surfaces sericeous to canescent.

Inflorescences

1–3-flowered.

1–3(–5)-flowered.

Peduncles

ascending or reflexed, upturned, 3–25 mm, shorter to longer than leaves (often elongated in fruit);

bract absent or 1–3-foliolate, usually subtending umbel.

ascending or spreading, (2–)10–20 mm, shorter to longer than leaves;

bract absent (reduced to stipules) or unifoliolate, distal.

Flowers

(5–)6–10(–12) mm;

calyx 3–5.5 mm, tube ± sparsely strigillose or glabrous, lobes subulate;

corolla yellow (sometimes banner orangish abaxially), turning orange or reddish, claws ± equaling calyx tube, banner implicate-ascending or remaining closely implicate, wings longer than keel;

style corneously thickened, marked by color, straight, puberulent or glabrous.

5–12 mm;

calyx 4–7 mm, tube villous, lobes subulate;

corolla yellow, turning orange-red, claws ± equaling calyx tube, banner ascending to 45°, wings unequal, longer than keel;

style slightly curved or straight, glabrous or finely puberulent.

Legumes

persistent, exserted, erect or divergent, brown, straight to ± curved distally, compressed, not or slightly constricted, incompletely septate, linear-oblong, 10–35 × 2–3 mm, thinly leathery, apex initially short-beaked, dehiscent, smooth, margins smooth, thin, glabrous or ± strigose.

persistent, exserted, divergent, brown, straight or curved at tip, turgid, slightly constricted, imperfectly septate, linear-oblong to oblong, (10–)18–25 × 1.5–2 mm, leathery, apex short-beaked, dehiscent, smooth, margins smooth, strigillose to glabrate.

Seeds

5–10, greenish to brown, ± mottled, ovoid to cuboid or globose, rugulose or granular.

2–5+, olive brown, mottled, roundish-oblong to angled, smooth, 2-veined.

Acmispon strigosus

Acmispon argyraeus

Distribution
sw United States; n Mexico
[BONAP county map]
nw Mexico; California
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

D. Isely (1981) discussed the variation in Acmispon strigosus (as Lotus), distinguishing three varieties, two rather distinct but with intergrading phases, and one apparently more transitional, although distinct enough to be recognized. These varieties were only noted for California in recent treatments (for example, L. Brouillet 2012, following D. Isely 1993). Nonetheless, the typical (for example, excluding intergrading phases) varieties represent recognizable morphologies that also have geographic distinction. Specimens representing intergradation seem to occur throughout the range of overlap of these varieties, sometimes rendering determination difficult.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

Acmispon argyraeus (variety not specified) is reported to hybridize with A. rigidus (A. M. Ottley 1944).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Herbs evidently hirsute, especially at apex, greenish.
var. hirtellus
1. Herbs usually strigillose or canescent-tomentose to scantly pubescent, rarely strigillose, green or cinereous.
→ 2
2. Stems ascending or decumbent, not markedly succulent, usually appearing green, sometimes cinereous; leaflet blades linear-oblong, apices acute to obtuse; corolla opening; California.
var. strigosus
2. Stems prostrate, succulent, cinereous(–greenish); leaflet blades oblong, apices obtuse or truncate; corolla not opening; Arizona, California, Nevada.
var. tomentellus
1. Banner panduriform; pistils arched more than 90°, ovary-style junction defined by a line; leaflet blades oblanceolate to obovate, usually folded; legumes curved at tip.
var. multicaulis
1. Banner not panduriform; pistils arched less than 90°, ovary-style junction with a jog (style base broader than apex of ovary) or confluent; leaflet blades obovate, not folded; legumes usually straight.
→ 2
2. Herbs usually prostrate; stems branched, silvery-canescent; ovary-style junction with a jog; flowers 5–10 mm.
var. argyraeus
2. Herbs low-spreading or low-ascending; stems diffusely branched, green or canescent; ovary-style junction confluent; flowers (8–)10–12 mm.
var. notitius
Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Acmispon Fabaceae > subfam. Faboideae > Acmispon
Sibling taxa
A. americanus, A. argophyllus, A. argyraeus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. denticulatus, A. glaber, A. grandiflorus, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. tomentosus, A. utahensis, A. wrangelianus, A. wrightii
A. americanus, A. argophyllus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. denticulatus, A. glaber, A. grandiflorus, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. strigosus, A. tomentosus, A. utahensis, A. wrangelianus, A. wrightii
Subordinate taxa
A. strigosus var. hirtellus, A. strigosus var. strigosus, A. strigosus var. tomentellus
A. argyraeus var. argyraeus, A. argyraeus var. multicaulis, A. argyraeus var. notitius
Synonyms Hosackia strigosa, Anisolotus strigosus, Lotus strigosus, Ottleya strigosa Hosackia argyraea, Anisolotus argyraeus, Lotus argyraeus, Ottleya argyraea
Name authority (Nuttall) Brouillet: J. Bot. Res. Inst. Texas 2: 392. (2008) (Greene) Brouillet: J. Bot. Res. Inst. Texas 2: 389. (2008) — (as argyreus)
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