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strigose lotus

American bird's-foot trefoil, American deervetch, deer-vetch, prairie trefoil, Spanish clover, Spanish lotus

Habit Herbs, annual, usually mat-forming, sometimes cespitose (ascending and bushy), green to grayish, 0.3–5 dm, not or ± fleshy, strigillose, hirsute, canescent-tomentose, or scantily pubescent; taprooted. Herbs, annual, solitary or cespitose, green, 0.5–6(–12) dm, not fleshy, glabrate to ± pilose; taprooted.
Stems

1–20+, procumbent or decumbent to ascending, branched basally, herbaceous, slender, leafy.

1–10+, erect to procumbent, branched proximally, herbaceous, leafy.

Leaves

irregularly pinnate;

stipules glandlike;

subsessile;

rachis 3–20 mm, flattened;

leaflets 4–10, blades unequal, obovate to oblanceolate to linear-oblong or oblong, apex acute to obtuse or truncate, surfaces ± densely tomentose to glabrate.

pinnate;

stipules glandlike;

subsessile to short-petiolate;

rachis 0.5–3(–4) mm, not flattened;

leaflets (1–)3(–5), blades obovate, ovate, or elliptic to lanceolate, margins denticulate or entire, apex acute, surfaces sparsely appressed-hairy.

Inflorescences

1–3-flowered.

1(or 2)-flowered.

Peduncles

ascending or reflexed, upturned, 3–25 mm, shorter to longer than leaves (often elongated in fruit);

bract absent or 1–3-foliolate, usually subtending umbel.

in distal axils, ascending, 3–25 mm, longer than leaves;

bract unifoliolate, distal.

Flowers

(5–)6–10(–12) mm;

calyx 3–5.5 mm, tube ± sparsely strigillose or glabrous, lobes subulate;

corolla yellow (sometimes banner orangish abaxially), turning orange or reddish, claws ± equaling calyx tube, banner implicate-ascending or remaining closely implicate, wings longer than keel;

style corneously thickened, marked by color, straight, puberulent or glabrous.

(4–)5–9 mm;

calyx (2.5–)3–6.5 mm, tube pilose, lobes linear-lanceolate, accrescent;

corolla whitish or cream, turning pink or salmon, banner rose-striate, keel tip yellow, claws shorter than calyx tube, banner implicate-ascending to erect, wings ± equaling keel;

style abruptly angled upward 45°, glabrous.

Legumes

persistent, exserted, erect or divergent, brown, straight to ± curved distally, compressed, not or slightly constricted, incompletely septate, linear-oblong, 10–35 × 2–3 mm, thinly leathery, apex initially short-beaked, dehiscent, smooth, margins smooth, thin, glabrous or ± strigose.

persistent, exserted, spreading or pendent, brown, straight or ± curved, compressed, slightly constricted, not septate, linear-oblong to narrowly oblong, (15–)20–30(–40) × (1.5–)2–2.5 mm, thinly leathery, apex abruptly downward angled and curved, dehiscent, smooth, margins smooth, thickened, glabrous.

Seeds

5–10, greenish to brown, ± mottled, ovoid to cuboid or globose, rugulose or granular.

(3–)5–8, olive to light brown, often black-mottled, oblong, smooth.

Acmispon strigosus

Acmispon americanus

Distribution
sw United States; n Mexico
[BONAP county map]
North America; nw Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

D. Isely (1981) discussed the variation in Acmispon strigosus (as Lotus), distinguishing three varieties, two rather distinct but with intergrading phases, and one apparently more transitional, although distinct enough to be recognized. These varieties were only noted for California in recent treatments (for example, L. Brouillet 2012, following D. Isely 1993). Nonetheless, the typical (for example, excluding intergrading phases) varieties represent recognizable morphologies that also have geographic distinction. Specimens representing intergradation seem to occur throughout the range of overlap of these varieties, sometimes rendering determination difficult.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Herbs evidently hirsute, especially at apex, greenish.
var. hirtellus
1. Herbs usually strigillose or canescent-tomentose to scantly pubescent, rarely strigillose, green or cinereous.
→ 2
2. Stems ascending or decumbent, not markedly succulent, usually appearing green, sometimes cinereous; leaflet blades linear-oblong, apices acute to obtuse; corolla opening; California.
var. strigosus
2. Stems prostrate, succulent, cinereous(–greenish); leaflet blades oblong, apices obtuse or truncate; corolla not opening; Arizona, California, Nevada.
var. tomentellus
1. Herbs usually conspicuously pilose, sometimes glabrate; leaflet blades: length 3–3.5 times width.
var. americanus
1. Herbs glabrate; leaflet blades: length 4–5 times width.
var. helleri
Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Acmispon Fabaceae > subfam. Faboideae > Acmispon
Sibling taxa
A. americanus, A. argophyllus, A. argyraeus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. denticulatus, A. glaber, A. grandiflorus, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. tomentosus, A. utahensis, A. wrangelianus, A. wrightii
A. argophyllus, A. argyraeus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. denticulatus, A. glaber, A. grandiflorus, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. strigosus, A. tomentosus, A. utahensis, A. wrangelianus, A. wrightii
Subordinate taxa
A. strigosus var. hirtellus, A. strigosus var. strigosus, A. strigosus var. tomentellus
A. americanus var. americanus, A. americanus var. helleri
Synonyms Hosackia strigosa, Anisolotus strigosus, Lotus strigosus, Ottleya strigosa Trigonella americana, Hosackia americana
Name authority (Nuttall) Brouillet: J. Bot. Res. Inst. Texas 2: 392. (2008) (Nuttall) Rydberg: Bull. Torrey Bot. Club 40: 45. (1913)
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