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chaparral bird's-foot trefoil, large leaf lotus, large-flower lotus

strigose lotus

Habit Herbs, perennial, cespitose, sometimes robust, grayish or green, 1–4(–15) dm, not fleshy, ± densely puberulent or strigillose; rhizomatous, woody based. Herbs, annual, usually mat-forming, sometimes cespitose (ascending and bushy), green to grayish, 0.3–5 dm, not or ± fleshy, strigillose, hirsute, canescent-tomentose, or scantily pubescent; taprooted.
Stems

1–5+, decumbent to erect, branched, herbaceous, often striate, leafy.

1–20+, procumbent or decumbent to ascending, branched basally, herbaceous, slender, leafy.

Leaves

irregularly pinnate;

stipules glandlike, conic; petiolate or sessile;

rachis 2–3.5(–5.5) mm, not flattened;

leaflets 7–9(–12), blades usually elliptic to obovate, sometimes ovate, apex acute to obtuse, surfaces sparsely to densely puberulent or villosulous to strigillose.

irregularly pinnate;

stipules glandlike;

subsessile;

rachis 3–20 mm, flattened;

leaflets 4–10, blades unequal, obovate to oblanceolate to linear-oblong or oblong, apex acute to obtuse or truncate, surfaces ± densely tomentose to glabrate.

Inflorescences

3–9(–11)-flowered.

1–3-flowered.

Peduncles

ascending or spreading, 10–80 mm, longer than leaves;

bract 1(–3)-foliolate, distal.

ascending or reflexed, upturned, 3–25 mm, shorter to longer than leaves (often elongated in fruit);

bract absent or 1–3-foliolate, usually subtending umbel.

Flowers

12–25 mm;

calyx (4.5–)5.5–10 mm, tube villosulous, lobes subulate;

corolla greenish white, white or yellow, fading to rose or reddish, claws shorter than calyx tube, banner ascending 45–90°, wings longer than banner and keel;

style nearly straight or basally curved, glabrous.

(5–)6–10(–12) mm;

calyx 3–5.5 mm, tube ± sparsely strigillose or glabrous, lobes subulate;

corolla yellow (sometimes banner orangish abaxially), turning orange or reddish, claws ± equaling calyx tube, banner implicate-ascending or remaining closely implicate, wings longer than keel;

style corneously thickened, marked by color, straight, puberulent or glabrous.

Legumes

persistent, exserted, brown, linear-oblong, straight, turgid, sometimes slightly constricted, incompletely septate, 25–42(–70) × 2–3 mm, leathery, apex short hook-beaked, dehiscent, smooth, margins smooth, thickened, glabrate.

persistent, exserted, erect or divergent, brown, straight to ± curved distally, compressed, not or slightly constricted, incompletely septate, linear-oblong, 10–35 × 2–3 mm, thinly leathery, apex initially short-beaked, dehiscent, smooth, margins smooth, thin, glabrous or ± strigose.

Seeds

5–9, olive to reddish brown, mottled, broadly ovoid, smooth.

5–10, greenish to brown, ± mottled, ovoid to cuboid or globose, rugulose or granular.

Acmispon grandiflorus

Acmispon strigosus

Distribution
nw Mexico; California
[WildflowerSearch map]
[BONAP county map]
sw United States; n Mexico
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

D. Isely (1981) discussed the variation in Acmispon strigosus (as Lotus), distinguishing three varieties, two rather distinct but with intergrading phases, and one apparently more transitional, although distinct enough to be recognized. These varieties were only noted for California in recent treatments (for example, L. Brouillet 2012, following D. Isely 1993). Nonetheless, the typical (for example, excluding intergrading phases) varieties represent recognizable morphologies that also have geographic distinction. Specimens representing intergradation seem to occur throughout the range of overlap of these varieties, sometimes rendering determination difficult.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Herbs densely puberulent or villosulous, usually grayish; ovules 22–30.
var. grandiflorus
1. Herbs strigillose or puberulent, green; ovules ca. 45.
var. macranthus
1. Herbs evidently hirsute, especially at apex, greenish.
var. hirtellus
1. Herbs usually strigillose or canescent-tomentose to scantly pubescent, rarely strigillose, green or cinereous.
→ 2
2. Stems ascending or decumbent, not markedly succulent, usually appearing green, sometimes cinereous; leaflet blades linear-oblong, apices acute to obtuse; corolla opening; California.
var. strigosus
2. Stems prostrate, succulent, cinereous(–greenish); leaflet blades oblong, apices obtuse or truncate; corolla not opening; Arizona, California, Nevada.
var. tomentellus
Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Acmispon Fabaceae > subfam. Faboideae > Acmispon
Sibling taxa
A. americanus, A. argophyllus, A. argyraeus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. denticulatus, A. glaber, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. strigosus, A. tomentosus, A. utahensis, A. wrangelianus, A. wrightii
A. americanus, A. argophyllus, A. argyraeus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. denticulatus, A. glaber, A. grandiflorus, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. tomentosus, A. utahensis, A. wrangelianus, A. wrightii
Subordinate taxa
A. grandiflorus var. grandiflorus, A. grandiflorus var. macranthus
A. strigosus var. hirtellus, A. strigosus var. strigosus, A. strigosus var. tomentellus
Synonyms Hosackia grandiflora, Anisolotus grandiflorus, Lotus grandiflorus, Ottleya grandiflora Hosackia strigosa, Anisolotus strigosus, Lotus strigosus, Ottleya strigosa
Name authority (Bentham) Brouillet: J. Bot. Res. Inst. Texas 2: 390. (2008) (Nuttall) Brouillet: J. Bot. Res. Inst. Texas 2: 392. (2008)
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