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meadow birds-foot trefoil, meadow lotus, Mohave trefoil, riverbar bird's-foot-trefoil, riverbar lotus, riverbar trefoil, tooth lotus

Habit Herbs, annual, cespitose, often glaucous, 0.3–4 dm, not fleshy, glabrous or hirsute; taprooted. Herbs, perennial (often flowering first year, appearing annual), mat-forming, low, green, 3–10 dm, not fleshy, usually distally villous (young), sparsely hirsute to glabrate (mature); from woody caudices, taprooted.
Stems

1(–5), decumbent to erect, apically or basally coarse-branched, herbaceous, leafy.

1–10+, procumbent, much branched, wiry or ± woody, leafy.

Leaves

subpinnate, pinnate, or palmate;

stipules glandlike or absent; petiolate;

rachis 5–12 mm, flattened;

leaflets 2–4, often 1 or 2 on one side and 2 terminal, blades elliptic to obovate (lateral sometimes asymmetric), margins denticulate or entire, apex acute to obtuse, surfaces hirsute.

irregularly subpalmate;

stipules glandlike;

subessile to petiolate;

rachis 2–10 mm, sometimes flattened;

leaflets 4–6, blades ovate to obovate, apex ± obtuse to acute, surfaces sparsely to densely pilose.

Inflorescences

1 or 2-flowered.

3–8-flowered.

Peduncles

± sessile;

bract absent.

ascending, 1–5 mm, shorter than leaves;

bract unifoliolate, distal.

Flowers

5–8 mm;

calyx 3–5 mm, tube hirsute or glabrous, lobes subulate, ± denticulate;

corolla cream-white to pale yellow, banner purple-tinged, keel tip yellowish, claws shorter to slightly longer than calyx tube, banner ascending, wings ± equaling keel, with deep, triangular auricle;

style curved, glabrous.

4–6(–7) mm;

calyx 2–4 mm, tube usually villous, lobes setaceous;

corolla yellow to reddish, dark-tipped, claws shorter than or equaling calyx tube, banner implicate-ascending, wings equaling to longer than keel (and other petals);

style geniculate 90° or incurved, glabrous.

Legumes

persistent, solitary or paired, exserted, erect or spreading, tawny, straight, compressed, slightly constricted, not septate, widely oblong, 8–20 × 3 mm, leathery, apex abruptly downward angled and curved, dehiscent, smooth, margins often undulate-verrucose, strigose or glabrous.

deciduous with calyx, exserted, ascending to divergent, tawny, straight to strongly arched, turgid, not constricted, not septate, linear-oblong, 3–5 × 1–2 mm, leathery, apex abruptly tapering to long, curved beak, indehiscent, smooth, margins indistinctly keeled, smooth, villous.

Seeds

(2 or)3(or 4), gray, faintly mottled, asymmetrically ± angular-obovoid, flattened, smooth.

1 or 2, olive green to brownish green, not mottled, straight to curved, narrowly ellipsoid, smooth.

2n

= 12.

Acmispon denticulatus

Acmispon tomentosus

Phenology Flowering spring–summer.
Habitat Grassy slopes, meadows, prairies, clearings, gravel bars, stream banks, vernal pools, pastures, grainfields, usually sandy soils, sometimes alkali, clay, or serpentine soils, roadsides.
Elevation 0–1900 m. (0–6200 ft.)
Distribution
from FNA
CA; ID; OR; UT; WA; BC
[WildflowerSearch map]
[BONAP county map]
nw Mexico; California
Discussion

Acmispon denticulatus occurs in California from the San Francisco Bay area, Sacramento Valley, and northern Sierra Nevada Foothills to the northwest, Cascade Range and Modoc Plateau, into adjacent southern Oregon (Siskiyou and Klamath regions), northward on both sides of the Cascade Range into southern British Columbia, with eastern outliers in southwestern Utah (Washington County), and in south-central Idaho (Lincoln County).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems usually villous distally; flowers 5–6(–7) mm; ovaries usually ± densely strigose.
var. tomentosus
1. Stems glabrate or villous distally; flowers 4–5 mm; ovaries usually sparsely strigose.
var. glabriusculus
Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Acmispon Fabaceae > subfam. Faboideae > Acmispon
Sibling taxa
A. americanus, A. argophyllus, A. argyraeus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. glaber, A. grandiflorus, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. strigosus, A. tomentosus, A. utahensis, A. wrangelianus, A. wrightii
A. americanus, A. argophyllus, A. argyraeus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. denticulatus, A. glaber, A. grandiflorus, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. strigosus, A. utahensis, A. wrangelianus, A. wrightii
Subordinate taxa
A. tomentosus var. glabriusculus, A. tomentosus var. tomentosus
Synonyms Hosackia denticulata, Anisolotus denticulatus, Lotus denticulatus Hosackia tomentosa
Name authority (Drew) D. D. Sokoloff: Ann. Bot. Fenn. 37: 130. (2000) (Hooker & Arnott) Govaerts: Skvortsovia 4(3): 76. (2018)
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