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meadow birds-foot trefoil, meadow lotus, Mohave trefoil, riverbar bird's-foot-trefoil, riverbar lotus, riverbar trefoil, tooth lotus

Habit Herbs, annual, cespitose, often glaucous, 0.3–4 dm, not fleshy, glabrous or hirsute; taprooted. Herbs, perennial, mat-forming or cespitose, diffuse, green, 0.5–4.5 dm, not fleshy, ± strigose to puberulent; from woody caudices.
Stems

1(–5), decumbent to erect, apically or basally coarse-branched, herbaceous, leafy.

1–10+, decumbent to ascending, branched, herbaceous, sometimes slender, leafy, base with or without persistent leaves.

Leaves

subpinnate, pinnate, or palmate;

stipules glandlike or absent; petiolate;

rachis 5–12 mm, flattened;

leaflets 2–4, often 1 or 2 on one side and 2 terminal, blades elliptic to obovate (lateral sometimes asymmetric), margins denticulate or entire, apex acute to obtuse, surfaces hirsute.

irregularly pinnate to subpalmate with a terminal trefoil, often dimorphic (proximal with smaller, rounder leaflets);

stipules glandlike;

subsessile to short-petiolate;

rachis (1.5–)5–10 mm, flattened;

leaflets (3 or)4 or 5(–7)[–13], blades obovate (widely obovate) to linear-oblanceolate [elliptic], apex emarginate or obtuse to acute, surfaces strigose to villous.

Inflorescences

1 or 2-flowered.

1–3(or 4)-flowered.

Peduncles

± sessile;

bract absent.

ascending, 10–70(–90) mm, longer than leaves;

bract absent or 1–3[–5]-foliolate, subtending umbel.

Flowers

5–8 mm;

calyx 3–5 mm, tube hirsute or glabrous, lobes subulate, ± denticulate;

corolla cream-white to pale yellow, banner purple-tinged, keel tip yellowish, claws shorter to slightly longer than calyx tube, banner ascending, wings ± equaling keel, with deep, triangular auricle;

style curved, glabrous.

10–16(–20) mm;

calyx 5–8.5[–10] mm, tube villous, lobes lanceolate to subulate;

corolla yellow, turning orange or red [banner red, keel and wings white], claws ± equaling calyx tube, banner ascending to 45°, wings longer than keel;

style straight, glabrous.

Legumes

persistent, solitary or paired, exserted, erect or spreading, tawny, straight, compressed, slightly constricted, not septate, widely oblong, 8–20 × 3 mm, leathery, apex abruptly downward angled and curved, dehiscent, smooth, margins often undulate-verrucose, strigose or glabrous.

persistent, exserted, erect to spreading, green or purplish brown [ashy], straight or ± curved, compressed, slightly constricted, imperfectly septate, linear-oblong, 17–35 × 2–3 mm, leathery, apex short-beaked, dehiscent, smooth, margins keeled, strigillose [glabrate].

Seeds

(2 or)3(or 4), gray, faintly mottled, asymmetrically ± angular-obovoid, flattened, smooth.

10–18, dark brown, ± mottled, oblong, dull, ± compressed, smooth.

2n

= 12.

= 14.

Acmispon denticulatus

Acmispon plebeius

Phenology Flowering spring–summer. Flowering spring or fall.
Habitat Grassy slopes, meadows, prairies, clearings, gravel bars, stream banks, vernal pools, pastures, grainfields, usually sandy soils, sometimes alkali, clay, or serpentine soils, roadsides. Sandy or gravelly, often dry, sometimes moist soils, on rock, flats, creekbeds, talus or colluvial slopes, rocky hillsides, ridges, canyons, grasslands, savannas, montane or desert scrub, cypress or juniper-pinyon or oak woodlands, oak-Douglas-fir-pine, ponderosa pine or pine-oak forests, washes, roadsides.
Elevation 0–1900 m. (0–6200 ft.) 1000–2700 m. (3300–8900 ft.)
Distribution
from FNA
CA; ID; OR; UT; WA; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM; NV; TX; UT; Mexico (Baja California, Baja California Sur, Chihuahua, Durango, Hidalgo, Jalisco, Puebla)
Discussion

Acmispon denticulatus occurs in California from the San Francisco Bay area, Sacramento Valley, and northern Sierra Nevada Foothills to the northwest, Cascade Range and Modoc Plateau, into adjacent southern Oregon (Siskiyou and Klamath regions), northward on both sides of the Cascade Range into southern British Columbia, with eastern outliers in southwestern Utah (Washington County), and in south-central Idaho (Lincoln County).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Acmispon plebeius occurs in the plateaus of southern Utah and into Lincoln County, Nevada, as well as in the plateaus and ranges of Arizona and New Mexico. It is reported to hybridize with A. neomexicanus and A. rigidus (A. M. Ottley 1944; D. Isely 1981), as well as A. wrightii (Isely).

Lotus nummulus Dayton is an illegitimate, superfluous name that pertains here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Acmispon Fabaceae > subfam. Faboideae > Acmispon
Sibling taxa
A. americanus, A. argophyllus, A. argyraeus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. glaber, A. grandiflorus, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. strigosus, A. tomentosus, A. utahensis, A. wrangelianus, A. wrightii
A. americanus, A. argophyllus, A. argyraeus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. denticulatus, A. glaber, A. grandiflorus, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. strigosus, A. tomentosus, A. utahensis, A. wrangelianus, A. wrightii
Synonyms Hosackia denticulata, Anisolotus denticulatus, Lotus denticulatus Hosackia plebeia, Anisolotus longebracteatus, A. nummularius, H. rigida var. nummularia, Lotus longebracteatus, L. nummularius, L. oroboides var. nanus, L. oroboides var. nummularius, L. oroboides var. plebeius, L. oroboides var. ramulosus, L. plebeius, L. ramulosus, Ottleya plebeia
Name authority (Drew) D. D. Sokoloff: Ann. Bot. Fenn. 37: 130. (2000) (Brandegee) Brouillet: Phytoneuron 2020-29: 2. (2020)
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