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meadow birds-foot trefoil, meadow lotus, Mohave trefoil, riverbar bird's-foot-trefoil, riverbar lotus, riverbar trefoil, tooth lotus

chaparral bird's-foot trefoil, large leaf lotus, large-flower lotus

Habit Herbs, annual, cespitose, often glaucous, 0.3–4 dm, not fleshy, glabrous or hirsute; taprooted. Herbs, perennial, cespitose, sometimes robust, grayish or green, 1–4(–15) dm, not fleshy, ± densely puberulent or strigillose; rhizomatous, woody based.
Stems

1(–5), decumbent to erect, apically or basally coarse-branched, herbaceous, leafy.

1–5+, decumbent to erect, branched, herbaceous, often striate, leafy.

Leaves

subpinnate, pinnate, or palmate;

stipules glandlike or absent; petiolate;

rachis 5–12 mm, flattened;

leaflets 2–4, often 1 or 2 on one side and 2 terminal, blades elliptic to obovate (lateral sometimes asymmetric), margins denticulate or entire, apex acute to obtuse, surfaces hirsute.

irregularly pinnate;

stipules glandlike, conic; petiolate or sessile;

rachis 2–3.5(–5.5) mm, not flattened;

leaflets 7–9(–12), blades usually elliptic to obovate, sometimes ovate, apex acute to obtuse, surfaces sparsely to densely puberulent or villosulous to strigillose.

Inflorescences

1 or 2-flowered.

3–9(–11)-flowered.

Peduncles

± sessile;

bract absent.

ascending or spreading, 10–80 mm, longer than leaves;

bract 1(–3)-foliolate, distal.

Flowers

5–8 mm;

calyx 3–5 mm, tube hirsute or glabrous, lobes subulate, ± denticulate;

corolla cream-white to pale yellow, banner purple-tinged, keel tip yellowish, claws shorter to slightly longer than calyx tube, banner ascending, wings ± equaling keel, with deep, triangular auricle;

style curved, glabrous.

12–25 mm;

calyx (4.5–)5.5–10 mm, tube villosulous, lobes subulate;

corolla greenish white, white or yellow, fading to rose or reddish, claws shorter than calyx tube, banner ascending 45–90°, wings longer than banner and keel;

style nearly straight or basally curved, glabrous.

Legumes

persistent, solitary or paired, exserted, erect or spreading, tawny, straight, compressed, slightly constricted, not septate, widely oblong, 8–20 × 3 mm, leathery, apex abruptly downward angled and curved, dehiscent, smooth, margins often undulate-verrucose, strigose or glabrous.

persistent, exserted, brown, linear-oblong, straight, turgid, sometimes slightly constricted, incompletely septate, 25–42(–70) × 2–3 mm, leathery, apex short hook-beaked, dehiscent, smooth, margins smooth, thickened, glabrate.

Seeds

(2 or)3(or 4), gray, faintly mottled, asymmetrically ± angular-obovoid, flattened, smooth.

5–9, olive to reddish brown, mottled, broadly ovoid, smooth.

2n

= 12.

Acmispon denticulatus

Acmispon grandiflorus

Phenology Flowering spring–summer.
Habitat Grassy slopes, meadows, prairies, clearings, gravel bars, stream banks, vernal pools, pastures, grainfields, usually sandy soils, sometimes alkali, clay, or serpentine soils, roadsides.
Elevation 0–1900 m. (0–6200 ft.)
Distribution
from FNA
CA; ID; OR; UT; WA; BC
[WildflowerSearch map]
[BONAP county map]
nw Mexico; California
[WildflowerSearch map]
[BONAP county map]
Discussion

Acmispon denticulatus occurs in California from the San Francisco Bay area, Sacramento Valley, and northern Sierra Nevada Foothills to the northwest, Cascade Range and Modoc Plateau, into adjacent southern Oregon (Siskiyou and Klamath regions), northward on both sides of the Cascade Range into southern British Columbia, with eastern outliers in southwestern Utah (Washington County), and in south-central Idaho (Lincoln County).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Herbs densely puberulent or villosulous, usually grayish; ovules 22–30.
var. grandiflorus
1. Herbs strigillose or puberulent, green; ovules ca. 45.
var. macranthus
Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Acmispon Fabaceae > subfam. Faboideae > Acmispon
Sibling taxa
A. americanus, A. argophyllus, A. argyraeus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. glaber, A. grandiflorus, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. strigosus, A. tomentosus, A. utahensis, A. wrangelianus, A. wrightii
A. americanus, A. argophyllus, A. argyraeus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. denticulatus, A. glaber, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. strigosus, A. tomentosus, A. utahensis, A. wrangelianus, A. wrightii
Subordinate taxa
A. grandiflorus var. grandiflorus, A. grandiflorus var. macranthus
Synonyms Hosackia denticulata, Anisolotus denticulatus, Lotus denticulatus Hosackia grandiflora, Anisolotus grandiflorus, Lotus grandiflorus, Ottleya grandiflora
Name authority (Drew) D. D. Sokoloff: Ann. Bot. Fenn. 37: 130. (2000) (Bentham) Brouillet: J. Bot. Res. Inst. Texas 2: 390. (2008)
Web links