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meadow birds-foot trefoil, meadow lotus, Mohave trefoil, riverbar bird's-foot-trefoil, riverbar lotus, riverbar trefoil, tooth lotus

canyon bird's-foot trefoil

Habit Herbs, annual, cespitose, often glaucous, 0.3–4 dm, not fleshy, glabrous or hirsute; taprooted. Herbs, perennial, mat-forming, silvery or gray, 1–3 dm, not fleshy, sericeous to canescent; from woody caudices.
Stems

1(–5), decumbent to erect, apically or basally coarse-branched, herbaceous, leafy.

1–20+, procumbent or decumbent to diffusely ascending, branched, herbaceous, slender, leafy.

Leaves

subpinnate, pinnate, or palmate;

stipules glandlike or absent; petiolate;

rachis 5–12 mm, flattened;

leaflets 2–4, often 1 or 2 on one side and 2 terminal, blades elliptic to obovate (lateral sometimes asymmetric), margins denticulate or entire, apex acute to obtuse, surfaces hirsute.

palmate or irregularly subpinnate;

stipules glandlike;

subsessile to short-petiolate;

rachis 1–4 mm, ± flattened;

leaflets 3–5, blades obovate to oblanceolate, apex usually obtuse, sometimes acute, surfaces sericeous to canescent.

Inflorescences

1 or 2-flowered.

1–3(–5)-flowered.

Peduncles

± sessile;

bract absent.

ascending or spreading, (2–)10–20 mm, shorter to longer than leaves;

bract absent (reduced to stipules) or unifoliolate, distal.

Flowers

5–8 mm;

calyx 3–5 mm, tube hirsute or glabrous, lobes subulate, ± denticulate;

corolla cream-white to pale yellow, banner purple-tinged, keel tip yellowish, claws shorter to slightly longer than calyx tube, banner ascending, wings ± equaling keel, with deep, triangular auricle;

style curved, glabrous.

5–12 mm;

calyx 4–7 mm, tube villous, lobes subulate;

corolla yellow, turning orange-red, claws ± equaling calyx tube, banner ascending to 45°, wings unequal, longer than keel;

style slightly curved or straight, glabrous or finely puberulent.

Legumes

persistent, solitary or paired, exserted, erect or spreading, tawny, straight, compressed, slightly constricted, not septate, widely oblong, 8–20 × 3 mm, leathery, apex abruptly downward angled and curved, dehiscent, smooth, margins often undulate-verrucose, strigose or glabrous.

persistent, exserted, divergent, brown, straight or curved at tip, turgid, slightly constricted, imperfectly septate, linear-oblong to oblong, (10–)18–25 × 1.5–2 mm, leathery, apex short-beaked, dehiscent, smooth, margins smooth, strigillose to glabrate.

Seeds

(2 or)3(or 4), gray, faintly mottled, asymmetrically ± angular-obovoid, flattened, smooth.

2–5+, olive brown, mottled, roundish-oblong to angled, smooth, 2-veined.

2n

= 12.

Acmispon denticulatus

Acmispon argyraeus

Phenology Flowering spring–summer.
Habitat Grassy slopes, meadows, prairies, clearings, gravel bars, stream banks, vernal pools, pastures, grainfields, usually sandy soils, sometimes alkali, clay, or serpentine soils, roadsides.
Elevation 0–1900 m. (0–6200 ft.)
Distribution
from FNA
CA; ID; OR; UT; WA; BC
[WildflowerSearch map]
[BONAP county map]
nw Mexico; California
[WildflowerSearch map]
[BONAP county map]
Discussion

Acmispon denticulatus occurs in California from the San Francisco Bay area, Sacramento Valley, and northern Sierra Nevada Foothills to the northwest, Cascade Range and Modoc Plateau, into adjacent southern Oregon (Siskiyou and Klamath regions), northward on both sides of the Cascade Range into southern British Columbia, with eastern outliers in southwestern Utah (Washington County), and in south-central Idaho (Lincoln County).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

Acmispon argyraeus (variety not specified) is reported to hybridize with A. rigidus (A. M. Ottley 1944).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Banner panduriform; pistils arched more than 90°, ovary-style junction defined by a line; leaflet blades oblanceolate to obovate, usually folded; legumes curved at tip.
var. multicaulis
1. Banner not panduriform; pistils arched less than 90°, ovary-style junction with a jog (style base broader than apex of ovary) or confluent; leaflet blades obovate, not folded; legumes usually straight.
→ 2
2. Herbs usually prostrate; stems branched, silvery-canescent; ovary-style junction with a jog; flowers 5–10 mm.
var. argyraeus
2. Herbs low-spreading or low-ascending; stems diffusely branched, green or canescent; ovary-style junction confluent; flowers (8–)10–12 mm.
var. notitius
Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Acmispon Fabaceae > subfam. Faboideae > Acmispon
Sibling taxa
A. americanus, A. argophyllus, A. argyraeus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. glaber, A. grandiflorus, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. strigosus, A. tomentosus, A. utahensis, A. wrangelianus, A. wrightii
A. americanus, A. argophyllus, A. brachycarpus, A. cytisoides, A. decumbens, A. dendroideus, A. denticulatus, A. glaber, A. grandiflorus, A. haydonii, A. intricatus, A. junceus, A. maritimus, A. mearnsii, A. micranthus, A. neomexicanus, A. parviflorus, A. plebeius, A. procumbens, A. prostratus, A. rigidus, A. rubriflorus, A. strigosus, A. tomentosus, A. utahensis, A. wrangelianus, A. wrightii
Subordinate taxa
A. argyraeus var. argyraeus, A. argyraeus var. multicaulis, A. argyraeus var. notitius
Synonyms Hosackia denticulata, Anisolotus denticulatus, Lotus denticulatus Hosackia argyraea, Anisolotus argyraeus, Lotus argyraeus, Ottleya argyraea
Name authority (Drew) D. D. Sokoloff: Ann. Bot. Fenn. 37: 130. (2000) (Greene) Brouillet: J. Bot. Res. Inst. Texas 2: 389. (2008) — (as argyreus)
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