Achnatherum thurberianum |
Achnatherum hymenoides |
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Thurber's needlegrass, Thurber's rice grass |
Indian rice grass, sand needlegrass, sand ricegrass |
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Habit | Plants cespitose, not rhizomatous. | Plants tightly cespitose, not rhizomatous. |
Culms | 30-75 cm tall, 0.5-1.7 mm thick, internodes pubescent or glabrous, pubescence more common on the lower internodes, particularly just below the nodes; nodes 2-3, lower nodes retrorsely pubescent, upper nodes glabrous or pubescent. |
25-70 cm tall, 0.7-1.3 mm thick, glabrous or partly scabridulous; nodes 3-4. |
Sheaths | glabrous or scabridulous, sometimes puberulent on the distal margins, hairs to 0.8 mm; collars glabrous, sometimes with tufts of hair on the sides, hairs to 1 mm; basal ligules 1.5-4 mm, hyaline, glabrous, acute; upper ligules to 2 mm; blades usually convolute, 0.1-1 mm in diameter, abaxial surfaces smooth or scabridulous, adaxial surfaces pubescent. |
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Panicles | 7-15 cm long, 0.5-2.5 cm wide, often included in the upper leaf sheaths at the start of anthesis; branches 1.5-6 cm, appressed to strongly ascending, with 1-6 spikelets. |
9-20 cm long, 8-14 cm wide; branches ascending to strongly divergent, longest branches 3-15 cm; pedicels paired, conspicuously divaricate, shorter pedicels in each pair usually at least 1/2 as long as the longer pedicels. |
Glumes | often purplish; lower glumes 10-15 mm long, 1.2-2 mm wide; upper glumes to 2 mm shorter; florets 6-9 mm long, 0.7-1.2 mm thick, fusiform, terete; calluses 0.9-1.5 mm, sharp; lemmas coriaceous, evenly pubescent or the back glabrate distally, hairs 0.5-0.8 mm, apices lobed on 1 margin, lobe about 0.1 mm long, thick, apical lemma hairs 0.5-0.8 mm; awns 32-56 mm, twice-geniculate, first 2 segments pilose, hairs 0.8-2 mm, terminal segment glabrous, often scabrous; paleas 4.6-6.1 mm, 3/4 - 9/10 as long as the lemmas, sparsely pubescent towards the base; anthers 2.5-3.5 mm, dehiscent, not penicillate. |
subequal, 5-9 mm long, 0.8-2 mm wide, saccate below, puberulent, hairs about 0.1 mm, tapering above midlength, apices acuminate; lower glumes 5-veined at the base, 3-veined at midlength; upper glumes 5-7-veined at the base; florets 3-4.5 mm long, 1-2 mm thick, obovoid; calluses 0.4-1 mm, sharp; lemmas indurate, densely and evenly pilose, hairs 2.5-6 mm, easily rubbed off, apices not lobed; awns 3-6 mm, rapidly deciduous, not geniculate, scabrous; paleas subequal to the lemmas in length and texture, glabrous, apices pinched; anthers 1.5-2 mm, penicillate, dehiscent, well-filled. |
Caryopses | 5-7 mm, fusiform. |
2-3 mm. |
Basal | sheaths glabrous, usually smooth, brown or gray-brown; collars glabrous, without tufts of hair at the sides; basal ligules 1.5-6 mm, hyaline, rounded to acute, lacerate; upper ligules to 8 mm, hyaline, acute, glabrous; blades 0.5-2 mm wide, convolute, abaxial surfaces scabrous, adaxial surfaces scabrous or hairy, hairs about 0.3 mm. |
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2n | =34. |
= 46, 48. |
Achnatherum thurberianum |
Achnatherum hymenoides |
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Distribution |
CA; CO; ID; MT; NV; OR; UT; WA; WY
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AR; AZ; CA; CO; ID; KS; MN; MT; ND; NE; NM; NV; OK; OR; SD; TX; UT; WA; WY; AB; BC; MB; SK; YT
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Discussion | Achnatherum thurberianum grows in canyons and foothills, primarily in sagebrush desert and juniper woodland associations, from Washington to southern Idaho and southwestern Montana and from California to Utah, at 900-3000 m. Its long ligules and pilose awns make it one of the easier North American species of Achnatherum to identify. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Achnatherum hymenoides grows in dry, well-drained soils, primarily in the western part of the Flora region and northern Mexico. Specimens from further east may be introduced; it is unknown whether they have persisted. The roots of A. hymenoides are often surrounded by a rhizosheath formed by mucilaginous secretions to which soil particles attach. This rhizosheath harbors nitrogen-fixing organisms that probably contribute to the success of the species as a colonizer. Native Americans used the seeds of Achnatherum hymenoides for food. It is also one of the most palatable native grasses for livestock. Several cultivars have been developed for use in restoration work, and it is becoming increasingly available for use as an ornamental. Achnatherum hymenoides forms natural hybrids with other members of the Stipeae. See discussion on p. 142. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 24, p. 125. | FNA vol. 24, p. 139. |
Parent taxa | Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum | Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum |
Sibling taxa | ||
Synonyms | Stipa thurberiana | Stipa hymenoides, Oryzopsis hymenoides |
Name authority | (Piper) Barkworth | (Roem. & Schult.) Barkworth |
Web links |
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