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sleepy grass

common western needlegrass, stiff needlegrass, velvet-leaf, western needle grass

Habit Plants cespitose, not rhizomatous. Plants tightly cespitose, not rhizomatous.
Culms

100-230 cm tall, 2-4.5 mm thick, mostly glabrous, often pubescent below the nodes, the pubescence antrorse or retrorse;

nodes 4-5.

14-120(180) cm tall, 0.3-2 mm thick, internodes glabrous or puberulent to densely pubescent;

nodes 2-4, glabrous or pubescent.

Panicles

15-30 cm long, 0.8-3.5 cm wide;

branches straight, appressed to ascending, lower branches 3-9 cm.

5-30 cm long, 0.5-1.5 cm wide;

branches appressed, straight, longest branches 1-7 cm.

Spikelets

appressed to the branches.

appressed to the branches.

Glumes

subequal, 9-11.5 mm long, 1-1.4 mm wide;

florets 5.9-8.5 mm long, 0.9-1.2 mm thick, fusiform, terete;

calluses 0.3-1 mm, blunt;

lemmas evenly hairy, hairs at midlength 0.3-0.8 mm, apical hairs to 1.5 mm;

awns 20-32 mm, persistent, twice-geniculate, scabridulous to scabrous, scabrules to about 0.1 mm, longest on the middle segment, terminal segment straight;

paleas 3.7-5.6 mm, 2/3 - 3/4 as long as the lemmas, hairy, hairs about 0.5 mm, not exceeding the apices, veins terminating below the apices, apices rounded;

anthers 4-5 mm, dehiscent, penicillate.

subequal, 9-15 mm long, 0.6-0.9 mm wide;

florets 5.5-7.5 mm long, 0.5-0.9 mm thick, fusiform, terete;

calluses 0.8-1.2 mm, sharp, dorsal boundary of the glabrous tip with the callus hairs narrowly acute;

lemmas evenly hair, hairs 0.5-1.5 mm at midlength, apical hairs somewhat longer than those below, sometimes similar in length to those at the base of the awns, sometimes longer, apical lobes 0.3-0.5 mm, membranous;

awns 15-55 mm, twice-geniculate, first 2 segments evidently hairy, terminal segment glabrous or partly to wholly pilose, sometimes scabrous;

paleas 2.2-3.5 mm, 2/5 – 3/5 as long as the lemmas, hairs at the tip usually shorter than 1 mm, frequently extending beyond the apices, apices rounded;

anthers 2.5-3.5 mm, dehiscent, not penicillate.

Caryopses

5-6 mm.

4-6 mm, fusiform.

Basal

sheaths mostly glabrous, margins usually ciliate distaily;

collars hairy, those of the flag leaves densely hairy, hairs 0.5-2 mm, sides glabrous;

basal ligules 1-2 mm;

upper ligules to 4 mm, truncate, rounded, or obtuse, glabrous;

blades 6-10 mm wide, glabrous, abaxial surfaces smooth, adaxial surfaces prominently ribbed, ribs scabrous.

sheaths glabrous or puberulent to densely pubescent, often ciliate at the throat;

collars often with tufts of hair at the sides;

ligules 0.2-1.5 mm, often ciliate;

blades 0.5-3 mm wide and flat, or convolute and 0.1-0.8 mm in diameter, lax to straight.

2n

= 64.

=36.

Achnatherum robustum

Achnatherum occidentale

Distribution
from FNA
AZ; CO; KS; NE; NM; SD; TX; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Achnatherum robustum grows on dry plains and hills, in open woods and forest clearings, and along roadsides, from Wyoming through Colorado to Arizona, New Mexico, and northern Mexico. Records from Kansas represent recent introductions; it is not clear whether the species has persisted there. Achnatherum robustum is sometimes confused with A. nelsonii subsp. dorei and Nassella viridula; it differs from both in the densely hairy collars of its flag leaves. Although not widely available, it has potential as an ornamental grass, particularly in arid regions with cold winters.

The English-language name refers to the effect some samples, particularly those from the Sacramento and Sierra Blanca mountains, New Mexico, have on livestock, especially horses and cattle. "Mildly poisoned animals are dejected, inactive, and withdrawn; severely poisoned animals lie on their sides in a profound slumber" (Cheeke and Shull 1985). The active ingredient is diacetone alcohol (Epstein et al 1964).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Achnatherum occidentale, which extends from British Columbia to California, Utah, and Colorado, varies considerably in pubescence and size. The three subspecies recognized here occasionally occur together.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Terminal awn segment usually pilose; culms 0.3-1 mm thick, glabrous even on the basal internodes; glumes often purplish
subsp. occidentale
1. Terminal awn segment usually scabrous or glabrous, occasionally pilose at the base; culms 0.5-2 mm thick; glumes usually green.
→ 2
2. First 2 awn segments scabrous or pilose with hairs of mixed lengths; apical lemma hairs longer than the basal awn hairs
subsp. californicum
2. First 2 awn segments pilose, the hairs gradually and evenly becoming shorter towards the first geniculation; apical lemma hairs similar in length to the basal awn hairs
subsp. pubescens
Source FNA vol. 24, p. 129. FNA vol. 24, p. 121.
Parent taxa Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum
Sibling taxa
A. aridum, A. arnowiae, A. contractum, A. coronatum, A. curvifolium, A. diegoense, A. eminens, A. hendersonii, A. hymenoides, A. latiglume, A. lemmonii, A. lettermanii, A. lobatum, A. nelsonii, A. nevadense, A. occidentale, A. parishii, A. perplexum, A. pinetorum, A. richardsonii, A. scribneri, A. splendens, A. stillmanii, A. swallenii, A. thurberianum, A. wallowaense, A. webberi, A. ×bloomeri
A. aridum, A. arnowiae, A. contractum, A. coronatum, A. curvifolium, A. diegoense, A. eminens, A. hendersonii, A. hymenoides, A. latiglume, A. lemmonii, A. lettermanii, A. lobatum, A. nelsonii, A. nevadense, A. parishii, A. perplexum, A. pinetorum, A. richardsonii, A. robustum, A. scribneri, A. splendens, A. stillmanii, A. swallenii, A. thurberianum, A. wallowaense, A. webberi, A. ×bloomeri
Subordinate taxa
A. occidentale subsp. californicum, A. occidentale subsp. occidentale, A. occidentale subsp. pubescens
Synonyms Stipa vaseyi, Stipa robusta Stipa occidentalis, Stipa occidentalis var. montana
Name authority (Vasey) Barkworth (Thurb.) Barkworth
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