Achnatherum robustum |
Achnatherum nelsonii |
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sleepy grass |
Columbia needle grass, Nelson achnatherum, Nelson's needlegrass |
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Habit | Plants cespitose, not rhizomatous. | Plants cespitose, not rhizomatous. | ||||
Culms | 100-230 cm tall, 2-4.5 mm thick, mostly glabrous, often pubescent below the nodes, the pubescence antrorse or retrorse; nodes 4-5. |
40-175 cm tall, 0.7-2.4 mm thick, lower cauline internodes usually glabrous, sometimes slightly pubescent below the lower nodes; nodes 2-5. |
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Panicles | 15-30 cm long, 0.8-3.5 cm wide; branches straight, appressed to ascending, lower branches 3-9 cm. |
9-36 cm long, 0.8-2 cm wide; branches ascending to appressed, straight. |
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Spikelets | appressed to the branches. |
appressed to the branches. |
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Glumes | subequal, 9-11.5 mm long, 1-1.4 mm wide; florets 5.9-8.5 mm long, 0.9-1.2 mm thick, fusiform, terete; calluses 0.3-1 mm, blunt; lemmas evenly hairy, hairs at midlength 0.3-0.8 mm, apical hairs to 1.5 mm; awns 20-32 mm, persistent, twice-geniculate, scabridulous to scabrous, scabrules to about 0.1 mm, longest on the middle segment, terminal segment straight; paleas 3.7-5.6 mm, 2/3 - 3/4 as long as the lemmas, hairy, hairs about 0.5 mm, not exceeding the apices, veins terminating below the apices, apices rounded; anthers 4-5 mm, dehiscent, penicillate. |
6-12.5 mm long, 0.7-1.1 mm wide; lower glumes exceeding the upper glumes by 0.2-0.8 mm; florets 4.5-7 mm long, 0.6-0.9 mm thick, fusiform; calluses 0.2-1 mm, blunt to sharp, dorsal boundary of the glabrous tip with the callus hairs almost straight to acute; lemmas evenly hairy, hairs at midlength 0.5-1 mm, hairs at the apices to 2 mm, erect to ascending, apical lobes 0.1-0.4 mm, membranous, flexible; awns 19-45 mm, persistent, twice-geniculate, first 2 segments scabrous or with hairs shorter than 0.5 mm, terminal segment straight; paleas 2-4 mm, 1/3 – 2/3 as long as the lemmas, pubescent, hairs usually not exceeding the apices, veins terminating before the apices, apices rounded; anthers 2-3.5 mm, dehiscent, not penicillate. |
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Caryopses | 5-6 mm. |
3-4 mm, fusiform. |
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Basal | sheaths mostly glabrous, margins usually ciliate distaily; collars hairy, those of the flag leaves densely hairy, hairs 0.5-2 mm, sides glabrous; basal ligules 1-2 mm; upper ligules to 4 mm, truncate, rounded, or obtuse, glabrous; blades 6-10 mm wide, glabrous, abaxial surfaces smooth, adaxial surfaces prominently ribbed, ribs scabrous. |
sheaths glabrous or sparsely to densely pubescent, margins sometimes ciliate; collars glabrous or somewhat pubescent, without tufts of hair on the sides, collars of the flag leaves glabrous or sparsely pubescent; basal ligules 0.2-0.7 mm, membranous, truncate to rounded, usually not ciliate; upper ligules 1-1.5 mm, acute; blades (0.5)1.2-5 mm wide. |
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2n | = 64. |
= 36, 44. |
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Achnatherum robustum |
Achnatherum nelsonii |
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Distribution |
AZ; CO; KS; NE; NM; SD; TX; WY
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AZ; CA; CO; ID; MT; NM; NV; OR; SD; TX; UT; WA; WY; AB; BC; SK; YT
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Discussion | Achnatherum robustum grows on dry plains and hills, in open woods and forest clearings, and along roadsides, from Wyoming through Colorado to Arizona, New Mexico, and northern Mexico. Records from Kansas represent recent introductions; it is not clear whether the species has persisted there. Achnatherum robustum is sometimes confused with A. nelsonii subsp. dorei and Nassella viridula; it differs from both in the densely hairy collars of its flag leaves. Although not widely available, it has potential as an ornamental grass, particularly in arid regions with cold winters. The English-language name refers to the effect some samples, particularly those from the Sacramento and Sierra Blanca mountains, New Mexico, have on livestock, especially horses and cattle. "Mildly poisoned animals are dejected, inactive, and withdrawn; severely poisoned animals lie on their sides in a profound slumber" (Cheeke and Shull 1985). The active ingredient is diacetone alcohol (Epstein et al 1964). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Achnatherum nelsonii grows in meadows and openings, from sagebrush steppe and pinyon-juniper woodlands to subalpine forests, at 500-3500 m. It flowers in late spring to early summer, differing in this respect from A. perplexum. It is sometimes sympatric with A. lettermanii, from which it differs in its shorter paleas and wider leaves, and its tendency to grow in deeper or less disturbed soils. It differs from A. lemmonii in having wider leaf blades, shorter paleas, and membranous lemma lobes, and from A. nevadense and A. occidentale in its scabrous awns and the truncate to acute boundary of the glabrous tip of the callus with the callus hairs. The two subspecies intergrade to some extent. There is also intergradation with Achnatherum occidentale, possibly as a result of hybridization and introgression. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24, p. 129. | FNA vol. 24, p. 123. | ||||
Parent taxa | Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum | Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Stipa vaseyi, Stipa robusta | Stipa williamsii, Stipa occidentalis var. nelsonii, Stipa nelsonii | ||||
Name authority | (Vasey) Barkworth | (Scribn.) Barkworth | ||||
Web links |
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