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Richardson needlegrass, Richardson's needlegrass, Richardson's rice grass, spreading needlegrass

Scribner needlegrass, Scribner's needlegrass

Habit Plants tightly cespitose, not rhizomatous. Plants cespitose, not rhizomatous.
Culms

30-100 cm tall, 1-1.5 mm thick, glabrous;

nodes usually 3.

25-90 cm tall, 0.5-1.6 mm thick, glabrous;

nodes 3.

Panicles

7-25 cm long, 7-15 cm wide;

branches divergent, flexuous, longest branches 7-10 cm, with the spikelets confined to the distal 1/4.

7-21 cm long, 0.5-1 cm wide;

branches appressed to ascending, straight.

Spikelets

pendulous.

Lower glumes

7.5-11 mm long, 0.9-1.2 mm wide;

upper glumes 2-3 mm shorter;

florets 5-6 mm long, 0.6-0.9 mm thick, fusiform, terete;

calluses 0.4-0.7 mm, blunt;

lemmas evenly hairy on the lower portion, often glabrate distally, body and apical hairs 0.2-0.5 mm, apical lobes not or scarcely developed, to 0.1 mm;

awns 15-25 mm, persistent, twice-geniculate, first 2 segments strigulose, hairs about 0.1 mm, terminal segment straight;

paleas 2.2-3.6 mm, 1/2 - 3/5 as long as the lemmas, pubescent, hairs not exceeding the apices, apices rounded;

anthers 2.5-3 mm, dehiscent, penicillate, hairs 0.1-0.5 mm.

10-17 mm long, 0.7-1.2 mm wide, exceeding the upper glumes by 2.5-4.5 mm, apices tapering, often slightly recurved;

florets 6-9.5 mm long, 0.6-1.1 mm thick, fusiform, terete, widest at or below midlength;

calluses 0.5-1.5 mm, sharp;

lemmas evenly hairy, hairs at midlength to 1 mm, apical hairs 2-3 mm, ascending, apical lobes 0.3-0.5 mm;

awns 13-25 mm, persistent, usually once-geniculate, first segment scabrous, terminal segment straight;

paleas 2.5-3.5 mm, 1/3 – 1/2 as long as the lemmas, pubescent, hairs not exceeding the apices, apices rounded;

anthers 3-5 mm, dehiscent, not penicillate.

Caryopses

3-4 mm, fusiform.

5-6 mm, fusiform.

Basal

sheaths glabrous, margins ciliolate;

collars glabrous, without tufts of hair on the sides;

ligules 0.1-0.5 mm, truncate, ciliolate;

blades 0.8-3 mm wide, convolute when dry, abaxial surfaces scabridulous, adaxial surfaces glabrous.

sheaths becoming flat and papery, margins ciliate distally;

collars glabrous, with tufts of hair on the sides, hairs on the basal leaves to 1.5 mm, hairs on the flag leaves 1-2.5 mm;

basal ligules 0.3-0.8 mm, truncate, erose, ciliate, cilia 0.2-0.4 mm;

upper ligules to 1.5 mm, asymmetric, obliquely truncate for most of their width, abruptly longer on 1 side;

blades to 30 cm long, 2-5 mm wide, flat or involute, long-tapering.

2n

= 44.

= 40.

Achnatherum richardsonii

Achnatherum scribneri

Distribution
from FNA
AK; CO; ID; MT; SD; WA; WY; AB; BC; MB; SK; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; NM; OK; TX; WY
[BONAP county map]
Discussion

Achnatherum richardsonii grows in open woodlands and grasslands,often on sand or gravel, from the Yukon Territory to Washington and Manitoba, and south in the Rocky Mountains through Montana and Wyoming to western South Dakota and northern Colorado. Its elevation range is 1000-3100 m. It is readily recognized by its combination of flexuous panicle branches, drooping spikelets, and straight distal awn segments. Scagel and Maze (1984) concluded that putative hybrids between A. richardsonii and A. nelsonii subsp. dorei were merely large plants of subsp. dorei that varied in the direction of A. richardsonii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Achnatherum scribneri grows on rocky slopes, in pinyon-juniper and ponderosa pine associations at 1500-2700 m, from southeastern Wyoming through Colorado to Arizona, New Mexico, western Oklahoma, and Texas, and in Capital Reef National Park, Utah. At present, the Utah population appears to be disjunct from the species' primary range; this may reflect a lack of collecting. Achnatherum scribneri is similar to A. parishii, A. robustum, A. perplexum, and A. lobatum, differing from all of them in its sharp calluses.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 133. FNA vol. 24, p. 135.
Parent taxa Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum
Sibling taxa
A. aridum, A. arnowiae, A. contractum, A. coronatum, A. curvifolium, A. diegoense, A. eminens, A. hendersonii, A. hymenoides, A. latiglume, A. lemmonii, A. lettermanii, A. lobatum, A. nelsonii, A. nevadense, A. occidentale, A. parishii, A. perplexum, A. pinetorum, A. robustum, A. scribneri, A. splendens, A. stillmanii, A. swallenii, A. thurberianum, A. wallowaense, A. webberi, A. ×bloomeri
A. aridum, A. arnowiae, A. contractum, A. coronatum, A. curvifolium, A. diegoense, A. eminens, A. hendersonii, A. hymenoides, A. latiglume, A. lemmonii, A. lettermanii, A. lobatum, A. nelsonii, A. nevadense, A. occidentale, A. parishii, A. perplexum, A. pinetorum, A. richardsonii, A. robustum, A. splendens, A. stillmanii, A. swallenii, A. thurberianum, A. wallowaense, A. webberi, A. ×bloomeri
Synonyms Stipa richardsonii Stipa scribneri
Name authority (Link) Barkworth (Vasey) Barkworth
Web links