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Richardson needlegrass, Richardson's needlegrass, Richardson's rice grass, spreading needlegrass

New Mexico needlegrass, perplexing needlegrass

Habit Plants tightly cespitose, not rhizomatous. Plants cespitose, not rhizomatous.
Culms

30-100 cm tall, 1-1.5 mm thick, glabrous;

nodes usually 3.

35-90 cm tall, 0.7-2.2 mm thick, lower internodes glabrous, puberulent to 5 mm below the nodes;

nodes 2-3.

Panicles

7-25 cm long, 7-15 cm wide;

branches divergent, flexuous, longest branches 7-10 cm, with the spikelets confined to the distal 1/4.

10-25 cm long, 0.5-1.5 cm wide;

branches ascending to appressed, straight.

Spikelets

pendulous.

appressed to the branches.

Glumes

unequal;

lower glumes 10-15 mm long, 0.5-1.1 mm wide, exceeding the upper glumes by 1-3(4) mm;

florets 5.5-11 mm long, 0.7-1 mm thick, fusiform, terete, widest at or below midlength;

calluses 0.4-0.6 mm, blunt;

lemmas evenly hairy, hairs at midlength about 1 mm, apical hairs 1-2 mm, ascending to divergent, apical lobes 0.2-0.5 mm, membranous, flexible;

awns 10-19 mm, persistent, once(twice)-geniculate, basal segments scabrous, terminal segments straight;

paleas 2.8-5.6 mm, 1/2 -2/3 as long as the lemmas, hairy, hairs not or scarcely exceeding the apices, veins terminating at or before the apices, apices acute to rounded;

anthers 2.5-4 mm, dehiscent, not penicillate.

Lower glumes

7.5-11 mm long, 0.9-1.2 mm wide;

upper glumes 2-3 mm shorter;

florets 5-6 mm long, 0.6-0.9 mm thick, fusiform, terete;

calluses 0.4-0.7 mm, blunt;

lemmas evenly hairy on the lower portion, often glabrate distally, body and apical hairs 0.2-0.5 mm, apical lobes not or scarcely developed, to 0.1 mm;

awns 15-25 mm, persistent, twice-geniculate, first 2 segments strigulose, hairs about 0.1 mm, terminal segment straight;

paleas 2.2-3.6 mm, 1/2 - 3/5 as long as the lemmas, pubescent, hairs not exceeding the apices, apices rounded;

anthers 2.5-3 mm, dehiscent, penicillate, hairs 0.1-0.5 mm.

Caryopses

3-4 mm, fusiform.

3-6 mm, fusiform.

Basal

sheaths glabrous, margins ciliolate;

collars glabrous, without tufts of hair on the sides;

ligules 0.1-0.5 mm, truncate, ciliolate;

blades 0.8-3 mm wide, convolute when dry, abaxial surfaces scabridulous, adaxial surfaces glabrous.

sheaths mostly glabrous, margins ciliolate distally;

collars glabrous, including the sides;

basal ligules 0.2-0.5 mm, truncate, ciliolate, cilia to 0.1 mm;

upper ligules 0.2-3.5 mm, rounded to acute;

blades to 30 cm long, 1-3 mm wide.

2n

= 44.

= unknown.

Achnatherum richardsonii

Achnatherum perplexum

Distribution
from FNA
AK; CO; ID; MT; SD; WA; WY; AB; BC; MB; SK; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM; TX; UT
[BONAP county map]
Discussion

Achnatherum richardsonii grows in open woodlands and grasslands,often on sand or gravel, from the Yukon Territory to Washington and Manitoba, and south in the Rocky Mountains through Montana and Wyoming to western South Dakota and northern Colorado. Its elevation range is 1000-3100 m. It is readily recognized by its combination of flexuous panicle branches, drooping spikelets, and straight distal awn segments. Scagel and Maze (1984) concluded that putative hybrids between A. richardsonii and A. nelsonii subsp. dorei were merely large plants of subsp. dorei that varied in the direction of A. richardsonii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Achnatherum perplexum grows on slopes in pinyon-pine associations of the southwestern United States and adjacent Mexico, at 1500-1700 m. It flowers in late summer to early fall. It has generally been confused with A. scribneri, A. nelsonii, and A. lobatum. It differs from A. scribneri in the glabrous collar margins of its basal leaves and its blunt calluses; from A. nelsonii and A. lettermanii in its unequal glumes; from A. lettermanii in its relatively short paleas; and from A. lobatum in its shorter lemma lobes and ascending to divergent apical lemma hairs.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 133. FNA vol. 24, p. 135.
Parent taxa Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum
Sibling taxa
A. aridum, A. arnowiae, A. contractum, A. coronatum, A. curvifolium, A. diegoense, A. eminens, A. hendersonii, A. hymenoides, A. latiglume, A. lemmonii, A. lettermanii, A. lobatum, A. nelsonii, A. nevadense, A. occidentale, A. parishii, A. perplexum, A. pinetorum, A. robustum, A. scribneri, A. splendens, A. stillmanii, A. swallenii, A. thurberianum, A. wallowaense, A. webberi, A. ×bloomeri
A. aridum, A. arnowiae, A. contractum, A. coronatum, A. curvifolium, A. diegoense, A. eminens, A. hendersonii, A. hymenoides, A. latiglume, A. lemmonii, A. lettermanii, A. lobatum, A. nelsonii, A. nevadense, A. occidentale, A. parishii, A. pinetorum, A. richardsonii, A. robustum, A. scribneri, A. splendens, A. stillmanii, A. swallenii, A. thurberianum, A. wallowaense, A. webberi, A. ×bloomeri
Synonyms Stipa richardsonii
Name authority (Link) Barkworth Hoge & Barkworth
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