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Richardson needlegrass, Richardson's needlegrass, Richardson's rice grass, spreading needlegrass

Letterman needlegrass, Letterman's needlegrass

Habit Plants tightly cespitose, not rhizomatous. Plants tightly cespitose, not rhizomatous.
Culms

30-100 cm tall, 1-1.5 mm thick, glabrous;

nodes usually 3.

15-90 cm tall, 0.5-0.8 mm thick, usually glabrous, sometimes puberulent to 5 mm below the lower nodes;

nodes 2-3.

Panicles

7-25 cm long, 7-15 cm wide;

branches divergent, flexuous, longest branches 7-10 cm, with the spikelets confined to the distal 1/4.

7-19 cm long, 0.5-1 cm wide;

branches straight, appressed to strongly ascending, longest branches 1.2-2.5 cm.

Spikelets

pendulous.

appressed to the branches.

Glumes

6.5-9 mm, subequal;

lower glumes 1(3)-veined;

upper glumes to 0.5 mm shorter than the lower glumes, 0.6-1 mm wide, 1-veined;

florets 4.5-6 mm long, 0.8-1 mm thick, fusiform, teret, widest below midlength;

calluses 0.4-1 mm, blunt;

lemmas evenly hairy, hairs at midlength about 0.5 mm, apical hairs 0.7-1.5(2) mm, apical lobes 0.3-0.8, membranous, flexible;

awns 12-25 mm, persistent, twice-geniculate, scabrous, terminal segment straight;

paleas 3-4 mm, 3/4 - 4/5(9/10) as long as the lemmas, veins terminating at or before the apices, apices round, flat, apical hairs 0.5-1 mm, extending beyond the palea body;

anthers 1.5-2 mm, dehiscent, not penicillate.

Lower glumes

7.5-11 mm long, 0.9-1.2 mm wide;

upper glumes 2-3 mm shorter;

florets 5-6 mm long, 0.6-0.9 mm thick, fusiform, terete;

calluses 0.4-0.7 mm, blunt;

lemmas evenly hairy on the lower portion, often glabrate distally, body and apical hairs 0.2-0.5 mm, apical lobes not or scarcely developed, to 0.1 mm;

awns 15-25 mm, persistent, twice-geniculate, first 2 segments strigulose, hairs about 0.1 mm, terminal segment straight;

paleas 2.2-3.6 mm, 1/2 - 3/5 as long as the lemmas, pubescent, hairs not exceeding the apices, apices rounded;

anthers 2.5-3 mm, dehiscent, penicillate, hairs 0.1-0.5 mm.

Caryopses

3-4 mm, fusiform.

about 4 mm, fusiform.

Basal

sheaths glabrous, margins ciliolate;

collars glabrous, without tufts of hair on the sides;

ligules 0.1-0.5 mm, truncate, ciliolate;

blades 0.8-3 mm wide, convolute when dry, abaxial surfaces scabridulous, adaxial surfaces glabrous.

sheaths smooth, glabrous, margins not ciliate;

collars, including the sides, glabrous or sparsely pubescent, collars of the flag leaves glabrous;

ligules 0.2-1.5(2) mm, without tufts of hair on the sides, truncate to rounded;

blades 0.5-2, wide, abaxial surfaces smooth to scabridulous, adaxial surfaces scabrous or puberulent.

2n

= 44.

= 32.

Achnatherum richardsonii

Achnatherum lettermanii

Distribution
from FNA
AK; CO; ID; MT; SD; WA; WY; AB; BC; MB; SK; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; MT; NM; NV; OR; UT; WY
[WildflowerSearch map]
[BONAP county map]
Discussion

Achnatherum richardsonii grows in open woodlands and grasslands,often on sand or gravel, from the Yukon Territory to Washington and Manitoba, and south in the Rocky Mountains through Montana and Wyoming to western South Dakota and northern Colorado. Its elevation range is 1000-3100 m. It is readily recognized by its combination of flexuous panicle branches, drooping spikelets, and straight distal awn segments. Scagel and Maze (1984) concluded that putative hybrids between A. richardsonii and A. nelsonii subsp. dorei were merely large plants of subsp. dorei that varied in the direction of A. richardsonii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Achnatherum lettermanii grows in meadows and on dry slopes, from sagebrush to subalpine habitats, at 1700-3400 m. Its range extends from Oregon and Montana to southern California, Arizona, and New Mexico; it is not known from Mexico. When sympatric with A. nelsonii, A. lettermanii tends to grow in shallower or more disturbed soils. It can be distinguished from that species by its generally finer leaves and more tightly cespitose growth habit, as well as its blunter calluses and longer paleas. Its relatively long paleas also distinguish A. lettermanii from A. perplexum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 133. FNA vol. 24, p. 118.
Parent taxa Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum
Sibling taxa
A. aridum, A. arnowiae, A. contractum, A. coronatum, A. curvifolium, A. diegoense, A. eminens, A. hendersonii, A. hymenoides, A. latiglume, A. lemmonii, A. lettermanii, A. lobatum, A. nelsonii, A. nevadense, A. occidentale, A. parishii, A. perplexum, A. pinetorum, A. robustum, A. scribneri, A. splendens, A. stillmanii, A. swallenii, A. thurberianum, A. wallowaense, A. webberi, A. ×bloomeri
A. aridum, A. arnowiae, A. contractum, A. coronatum, A. curvifolium, A. diegoense, A. eminens, A. hendersonii, A. hymenoides, A. latiglume, A. lemmonii, A. lobatum, A. nelsonii, A. nevadense, A. occidentale, A. parishii, A. perplexum, A. pinetorum, A. richardsonii, A. robustum, A. scribneri, A. splendens, A. stillmanii, A. swallenii, A. thurberianum, A. wallowaense, A. webberi, A. ×bloomeri
Synonyms Stipa richardsonii Stipa lettermanii
Name authority (Link) Barkworth (Vasey) Barkworth
Web links