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Richardson needlegrass, Richardson's needlegrass, Richardson's rice grass, spreading needlegrass

Southwestern needlegrass

Habit Plants tightly cespitose, not rhizomatous. Plants cespitose, shortly rhizomatous, bases knotty.
Culms

30-100 cm tall, 1-1.5 mm thick, glabrous;

nodes usually 3.

50-100 cm tall, 0.8-1.5 mm thick, glabrous;

nodes 2-3.

Panicles

7-25 cm long, 7-15 cm wide;

branches divergent, flexuous, longest branches 7-10 cm, with the spikelets confined to the distal 1/4.

20-55 cm long, 3-8 cm wide, open, often enclosed to midlength at anthesis;

lower branches 5-8 cm, ascending to divergent, flexuous.

Spikelets

pendulous.

Lower glumes

7.5-11 mm long, 0.9-1.2 mm wide;

upper glumes 2-3 mm shorter;

florets 5-6 mm long, 0.6-0.9 mm thick, fusiform, terete;

calluses 0.4-0.7 mm, blunt;

lemmas evenly hairy on the lower portion, often glabrate distally, body and apical hairs 0.2-0.5 mm, apical lobes not or scarcely developed, to 0.1 mm;

awns 15-25 mm, persistent, twice-geniculate, first 2 segments strigulose, hairs about 0.1 mm, terminal segment straight;

paleas 2.2-3.6 mm, 1/2 - 3/5 as long as the lemmas, pubescent, hairs not exceeding the apices, apices rounded;

anthers 2.5-3 mm, dehiscent, penicillate, hairs 0.1-0.5 mm.

5-12 mm long, 0.5-0.7 mm wide, 3-5-veined;

upper glumes 1-4 mm shorter, 3-veined;

florets 4-7.5 mm long, 0.5-0.9 mm thick, fusiform, terete;

calluses 1-2 mm, sharp;

lemmas evenly hairy, hairs 0.4-0.8 mm throughout, apical lobes not present;

awns 35-70 mm, persistent, twice-geniculate, first 2 segments scabrous, terminal segment flexuous;

paleas 1-2 mm, 1/3 – 1/2 as long as the lemmas, sparsely to moderately pubescent, apices rounded, flat;

anthers 3-3.5 mm, dehiscent, a few penicillate, hairs about 0.3 mm.

Caryopses

3-4 mm, fusiform.

about 4 mm, fusiform.

Basal

sheaths glabrous, margins ciliolate;

collars glabrous, without tufts of hair on the sides;

ligules 0.1-0.5 mm, truncate, ciliolate;

blades 0.8-3 mm wide, convolute when dry, abaxial surfaces scabridulous, adaxial surfaces glabrous.

sheaths mostly glabrous, ciliate on the margins;

collars glabrous on the back, usually with tufts of hair on the sides, hairs about 0.8 mm;

basal ligules 0.8-1.6 mm, membranous, glabrous, rounded to acute;

upper ligules to 4.5 mm, acute;

blades 0.7-3.5 mm wide, abaxial surfaces smooth to scabridulous, adaxial surfaces prominently ribbed, scabridulous or sparsely to densely pubescent, hairs about 0.1 mm.

2n

= 44.

= 44, 46.

Achnatherum richardsonii

Achnatherum eminens

Distribution
from FNA
AK; CO; ID; MT; SD; WA; WY; AB; BC; MB; SK; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM; TX
[WildflowerSearch map]
[BONAP county map]
Discussion

Achnatherum richardsonii grows in open woodlands and grasslands,often on sand or gravel, from the Yukon Territory to Washington and Manitoba, and south in the Rocky Mountains through Montana and Wyoming to western South Dakota and northern Colorado. Its elevation range is 1000-3100 m. It is readily recognized by its combination of flexuous panicle branches, drooping spikelets, and straight distal awn segments. Scagel and Maze (1984) concluded that putative hybrids between A. richardsonii and A. nelsonii subsp. dorei were merely large plants of subsp. dorei that varied in the direction of A. richardsonii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Achnatherum eminens grows on dry, rocky slopes and valleys in the mountains of the southwestern United States, primarily in desert scrub, at 600-2600 m. Its range extends into Mexico. It is easy to recognize because of its open panicle, flexuous branches, and flexuous awns. It is superficially similar to Nassella cernua, but differs in its longer, glabrous ligules, not or weakly overlapping lemma margins, pubescent paleas, and geographic distribution.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 133. FNA vol. 24, p. 133.
Parent taxa Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum
Sibling taxa
A. aridum, A. arnowiae, A. contractum, A. coronatum, A. curvifolium, A. diegoense, A. eminens, A. hendersonii, A. hymenoides, A. latiglume, A. lemmonii, A. lettermanii, A. lobatum, A. nelsonii, A. nevadense, A. occidentale, A. parishii, A. perplexum, A. pinetorum, A. robustum, A. scribneri, A. splendens, A. stillmanii, A. swallenii, A. thurberianum, A. wallowaense, A. webberi, A. ×bloomeri
A. aridum, A. arnowiae, A. contractum, A. coronatum, A. curvifolium, A. diegoense, A. hendersonii, A. hymenoides, A. latiglume, A. lemmonii, A. lettermanii, A. lobatum, A. nelsonii, A. nevadense, A. occidentale, A. parishii, A. perplexum, A. pinetorum, A. richardsonii, A. robustum, A. scribneri, A. splendens, A. stillmanii, A. swallenii, A. thurberianum, A. wallowaense, A. webberi, A. ×bloomeri
Synonyms Stipa richardsonii Stipa eminens
Name authority (Link) Barkworth (Cav.) Barkworth
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