The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

common western needlegrass, stiff needlegrass, velvet-leaf, western needle grass

Columbia needle grass, Nelson achnatherum, Nelson's needlegrass

Habit Plants tightly cespitose, not rhizomatous. Plants cespitose, not rhizomatous.
Culms

14-120(180) cm tall, 0.3-2 mm thick, internodes glabrous or puberulent to densely pubescent;

nodes 2-4, glabrous or pubescent.

40-175 cm tall, 0.7-2.4 mm thick, lower cauline internodes usually glabrous, sometimes slightly pubescent below the lower nodes;

nodes 2-5.

Panicles

5-30 cm long, 0.5-1.5 cm wide;

branches appressed, straight, longest branches 1-7 cm.

9-36 cm long, 0.8-2 cm wide;

branches ascending to appressed, straight.

Spikelets

appressed to the branches.

appressed to the branches.

Glumes

subequal, 9-15 mm long, 0.6-0.9 mm wide;

florets 5.5-7.5 mm long, 0.5-0.9 mm thick, fusiform, terete;

calluses 0.8-1.2 mm, sharp, dorsal boundary of the glabrous tip with the callus hairs narrowly acute;

lemmas evenly hair, hairs 0.5-1.5 mm at midlength, apical hairs somewhat longer than those below, sometimes similar in length to those at the base of the awns, sometimes longer, apical lobes 0.3-0.5 mm, membranous;

awns 15-55 mm, twice-geniculate, first 2 segments evidently hairy, terminal segment glabrous or partly to wholly pilose, sometimes scabrous;

paleas 2.2-3.5 mm, 2/5 – 3/5 as long as the lemmas, hairs at the tip usually shorter than 1 mm, frequently extending beyond the apices, apices rounded;

anthers 2.5-3.5 mm, dehiscent, not penicillate.

6-12.5 mm long, 0.7-1.1 mm wide;

lower glumes exceeding the upper glumes by 0.2-0.8 mm;

florets 4.5-7 mm long, 0.6-0.9 mm thick, fusiform;

calluses 0.2-1 mm, blunt to sharp, dorsal boundary of the glabrous tip with the callus hairs almost straight to acute;

lemmas evenly hairy, hairs at midlength 0.5-1 mm, hairs at the apices to 2 mm, erect to ascending, apical lobes 0.1-0.4 mm, membranous, flexible;

awns 19-45 mm, persistent, twice-geniculate, first 2 segments scabrous or with hairs shorter than 0.5 mm, terminal segment straight;

paleas 2-4 mm, 1/3 – 2/3 as long as the lemmas, pubescent, hairs usually not exceeding the apices, veins terminating before the apices, apices rounded;

anthers 2-3.5 mm, dehiscent, not penicillate.

Caryopses

4-6 mm, fusiform.

3-4 mm, fusiform.

Basal

sheaths glabrous or puberulent to densely pubescent, often ciliate at the throat;

collars often with tufts of hair at the sides;

ligules 0.2-1.5 mm, often ciliate;

blades 0.5-3 mm wide and flat, or convolute and 0.1-0.8 mm in diameter, lax to straight.

sheaths glabrous or sparsely to densely pubescent, margins sometimes ciliate;

collars glabrous or somewhat pubescent, without tufts of hair on the sides, collars of the flag leaves glabrous or sparsely pubescent;

basal ligules 0.2-0.7 mm, membranous, truncate to rounded, usually not ciliate;

upper ligules 1-1.5 mm, acute;

blades (0.5)1.2-5 mm wide.

2n

=36.

= 36, 44.

Achnatherum occidentale

Achnatherum nelsonii

Distribution
from FNA
AZ; CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; MT; NM; NV; OR; SD; TX; UT; WA; WY; AB; BC; SK; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Achnatherum occidentale, which extends from British Columbia to California, Utah, and Colorado, varies considerably in pubescence and size. The three subspecies recognized here occasionally occur together.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Achnatherum nelsonii grows in meadows and openings, from sagebrush steppe and pinyon-juniper woodlands to subalpine forests, at 500-3500 m. It flowers in late spring to early summer, differing in this respect from A. perplexum. It is sometimes sympatric with A. lettermanii, from which it differs in its shorter paleas and wider leaves, and its tendency to grow in deeper or less disturbed soils. It differs from A. lemmonii in having wider leaf blades, shorter paleas, and membranous lemma lobes, and from A. nevadense and A. occidentale in its scabrous awns and the truncate to acute boundary of the glabrous tip of the callus with the callus hairs.

The two subspecies intergrade to some extent. There is also intergradation with Achnatherum occidentale, possibly as a result of hybridization and introgression.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Terminal awn segment usually pilose; culms 0.3-1 mm thick, glabrous even on the basal internodes; glumes often purplish
subsp. occidentale
1. Terminal awn segment usually scabrous or glabrous, occasionally pilose at the base; culms 0.5-2 mm thick; glumes usually green.
→ 2
2. First 2 awn segments scabrous or pilose with hairs of mixed lengths; apical lemma hairs longer than the basal awn hairs
subsp. californicum
2. First 2 awn segments pilose, the hairs gradually and evenly becoming shorter towards the first geniculation; apical lemma hairs similar in length to the basal awn hairs
subsp. pubescens
1. Calluses blunt, dorsal boundary of the glabrous tip and the callus hairs almost straight to rounded; awns 19-31 mm long
subsp. dorei
1. Calluses sharp, dorsal boundary of the glabrous tip and the callus hairs acute; awns 19-45 mm long
subsp. nelsonii
Source FNA vol. 24, p. 121. FNA vol. 24, p. 123.
Parent taxa Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum
Sibling taxa
A. aridum, A. arnowiae, A. contractum, A. coronatum, A. curvifolium, A. diegoense, A. eminens, A. hendersonii, A. hymenoides, A. latiglume, A. lemmonii, A. lettermanii, A. lobatum, A. nelsonii, A. nevadense, A. parishii, A. perplexum, A. pinetorum, A. richardsonii, A. robustum, A. scribneri, A. splendens, A. stillmanii, A. swallenii, A. thurberianum, A. wallowaense, A. webberi, A. ×bloomeri
A. aridum, A. arnowiae, A. contractum, A. coronatum, A. curvifolium, A. diegoense, A. eminens, A. hendersonii, A. hymenoides, A. latiglume, A. lemmonii, A. lettermanii, A. lobatum, A. nevadense, A. occidentale, A. parishii, A. perplexum, A. pinetorum, A. richardsonii, A. robustum, A. scribneri, A. splendens, A. stillmanii, A. swallenii, A. thurberianum, A. wallowaense, A. webberi, A. ×bloomeri
Subordinate taxa
A. occidentale subsp. californicum, A. occidentale subsp. occidentale, A. occidentale subsp. pubescens
A. nelsonii subsp. dorei, A. nelsonii subsp. nelsonii
Synonyms Stipa occidentalis, Stipa occidentalis var. montana Stipa williamsii, Stipa occidentalis var. nelsonii, Stipa nelsonii
Name authority (Thurb.) Barkworth (Scribn.) Barkworth
Web links