Achnatherum occidentale |
Achnatherum arnowiae |
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common western needlegrass, stiff needlegrass, velvet-leaf, western needle grass |
Arnow's ricegrass |
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Habit | Plants tightly cespitose, not rhizomatous. | Plants tightly cespitose, not rhizomatous. | ||||||||
Culms | 14-120(180) cm tall, 0.3-2 mm thick, internodes glabrous or puberulent to densely pubescent; nodes 2-4, glabrous or pubescent. |
15-75 cm tall. |
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Sheaths | smooth, mostly glabrous, margins ciliate; collars glabrous, with or without tufts of hair at the sides, hairs 0.7-2 mm; ligules 1-4 mm, glabrous or sparsely hairy, acute, sometimes ciliate; blades 1-2 mm wide, involute, 0.5-1 mm in diameter, abaxial surfaces scabridulous or smooth, adaxial surfaces densely hairy, hairs about 0.2 mm. |
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Panicles | 5-30 cm long, 0.5-1.5 cm wide; branches appressed, straight, longest branches 1-7 cm. |
5-20 cm long, 0.5-2.8 cm wide, loosely contracted; branches strongly ascending, longest branches 0.5-2.5(5) cm. |
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Spikelets | appressed to the branches. |
evenly distributed along the branches; pedicels loosely appressed to the branches, paired, unequal, shorter pedicels in each pair usually less than 1/2 as long as the longer pedicels. |
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Glumes | subequal, 9-15 mm long, 0.6-0.9 mm wide; florets 5.5-7.5 mm long, 0.5-0.9 mm thick, fusiform, terete; calluses 0.8-1.2 mm, sharp, dorsal boundary of the glabrous tip with the callus hairs narrowly acute; lemmas evenly hair, hairs 0.5-1.5 mm at midlength, apical hairs somewhat longer than those below, sometimes similar in length to those at the base of the awns, sometimes longer, apical lobes 0.3-0.5 mm, membranous; awns 15-55 mm, twice-geniculate, first 2 segments evidently hairy, terminal segment glabrous or partly to wholly pilose, sometimes scabrous; paleas 2.2-3.5 mm, 2/5 – 3/5 as long as the lemmas, hairs at the tip usually shorter than 1 mm, frequently extending beyond the apices, apices rounded; anthers 2.5-3.5 mm, dehiscent, not penicillate. |
slightly unequal, saccate below, tapering from about midlength, veins and sometimes also the intercostal regions puberulent, hairs to 0.1 mm, apices acute to acuminate; lower glumes 5.1-6.1 mm; upper glumes 4.3-5.2 mm; florets 2.8-4.2 mm, ovoid; calluses 0.2-0.4 mm, acute; lemmas indurate, dark gray-brown, smooth, densely pilose, hairs at midlength and at the apices similar, 2-3 mm, easily rubbed off, apices not lobed; awns 3-4.4 mm, rapidly deciduous, not geniculate, scabrous; paleas similar to the lemmas in length and texture, glabrous, apices pinched; anthers about 1 mm, dehiscent, penicillate. |
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Caryopses | 4-6 mm, fusiform. |
1-1.7 mm long, 0.8-1 mm in diameter, globose to obovoid. |
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Basal | sheaths glabrous or puberulent to densely pubescent, often ciliate at the throat; collars often with tufts of hair at the sides; ligules 0.2-1.5 mm, often ciliate; blades 0.5-3 mm wide and flat, or convolute and 0.1-0.8 mm in diameter, lax to straight. |
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2n | =36. |
= unknown. |
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Achnatherum occidentale |
Achnatherum arnowiae |
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Distribution |
AZ; CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC
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Discussion | Achnatherum occidentale, which extends from British Columbia to California, Utah, and Colorado, varies considerably in pubescence and size. The three subspecies recognized here occasionally occur together. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Achnatherum arnowiae grows in pinyon-juniper, sagebrush, and mixed desert shrub communities in Utah, at 1400-2000 m. Welsh and Atwood (2003) state that specimens belonging to A. arnowiae are often filed as A. xbloomeri, and they suggest that this species may also be hybrid in origin, possibly with Hesperostipa comata as one of its parents. Another possibility is that it is a derivative of A. hymenoides that is adapted to particular soil types. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24, p. 121. | FNA vol. 24, p. 141. | ||||||||
Parent taxa | Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum | Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Stipa occidentalis, Stipa occidentalis var. montana | |||||||||
Name authority | (Thurb.) Barkworth | (S.L. Welsh & N.D Atwood) Barkworth | ||||||||
Web links |
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