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common western needlegrass, stiff needlegrass, velvet-leaf, western needle grass

Arnow's ricegrass

Habit Plants tightly cespitose, not rhizomatous. Plants tightly cespitose, not rhizomatous.
Culms

14-120(180) cm tall, 0.3-2 mm thick, internodes glabrous or puberulent to densely pubescent;

nodes 2-4, glabrous or pubescent.

15-75 cm tall.

Sheaths

smooth, mostly glabrous, margins ciliate;

collars glabrous, with or without tufts of hair at the sides, hairs 0.7-2 mm;

ligules 1-4 mm, glabrous or sparsely hairy, acute, sometimes ciliate;

blades 1-2 mm wide, involute, 0.5-1 mm in diameter, abaxial surfaces scabridulous or smooth, adaxial surfaces densely hairy, hairs about 0.2 mm.

Panicles

5-30 cm long, 0.5-1.5 cm wide;

branches appressed, straight, longest branches 1-7 cm.

5-20 cm long, 0.5-2.8 cm wide, loosely contracted;

branches strongly ascending, longest branches 0.5-2.5(5) cm.

Spikelets

appressed to the branches.

evenly distributed along the branches;

pedicels loosely appressed to the branches, paired, unequal, shorter pedicels in each pair usually less than 1/2 as long as the longer pedicels.

Glumes

subequal, 9-15 mm long, 0.6-0.9 mm wide;

florets 5.5-7.5 mm long, 0.5-0.9 mm thick, fusiform, terete;

calluses 0.8-1.2 mm, sharp, dorsal boundary of the glabrous tip with the callus hairs narrowly acute;

lemmas evenly hair, hairs 0.5-1.5 mm at midlength, apical hairs somewhat longer than those below, sometimes similar in length to those at the base of the awns, sometimes longer, apical lobes 0.3-0.5 mm, membranous;

awns 15-55 mm, twice-geniculate, first 2 segments evidently hairy, terminal segment glabrous or partly to wholly pilose, sometimes scabrous;

paleas 2.2-3.5 mm, 2/5 – 3/5 as long as the lemmas, hairs at the tip usually shorter than 1 mm, frequently extending beyond the apices, apices rounded;

anthers 2.5-3.5 mm, dehiscent, not penicillate.

slightly unequal, saccate below, tapering from about midlength, veins and sometimes also the intercostal regions puberulent, hairs to 0.1 mm, apices acute to acuminate;

lower glumes 5.1-6.1 mm;

upper glumes 4.3-5.2 mm;

florets 2.8-4.2 mm, ovoid;

calluses 0.2-0.4 mm, acute;

lemmas indurate, dark gray-brown, smooth, densely pilose, hairs at midlength and at the apices similar, 2-3 mm, easily rubbed off, apices not lobed;

awns 3-4.4 mm, rapidly deciduous, not geniculate, scabrous;

paleas similar to the lemmas in length and texture, glabrous, apices pinched;

anthers about 1 mm, dehiscent, penicillate.

Caryopses

4-6 mm, fusiform.

1-1.7 mm long, 0.8-1 mm in diameter, globose to obovoid.

Basal

sheaths glabrous or puberulent to densely pubescent, often ciliate at the throat;

collars often with tufts of hair at the sides;

ligules 0.2-1.5 mm, often ciliate;

blades 0.5-3 mm wide and flat, or convolute and 0.1-0.8 mm in diameter, lax to straight.

2n

=36.

= unknown.

Achnatherum occidentale

Achnatherum arnowiae

Distribution
from FNA
AZ; CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
[BONAP county map]
Discussion

Achnatherum occidentale, which extends from British Columbia to California, Utah, and Colorado, varies considerably in pubescence and size. The three subspecies recognized here occasionally occur together.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Achnatherum arnowiae grows in pinyon-juniper, sagebrush, and mixed desert shrub communities in Utah, at 1400-2000 m. Welsh and Atwood (2003) state that specimens belonging to A. arnowiae are often filed as A. xbloomeri, and they suggest that this species may also be hybrid in origin, possibly with Hesperostipa comata as one of its parents. Another possibility is that it is a derivative of A. hymenoides that is adapted to particular soil types.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Terminal awn segment usually pilose; culms 0.3-1 mm thick, glabrous even on the basal internodes; glumes often purplish
subsp. occidentale
1. Terminal awn segment usually scabrous or glabrous, occasionally pilose at the base; culms 0.5-2 mm thick; glumes usually green.
→ 2
2. First 2 awn segments scabrous or pilose with hairs of mixed lengths; apical lemma hairs longer than the basal awn hairs
subsp. californicum
2. First 2 awn segments pilose, the hairs gradually and evenly becoming shorter towards the first geniculation; apical lemma hairs similar in length to the basal awn hairs
subsp. pubescens
Source FNA vol. 24, p. 121. FNA vol. 24, p. 141.
Parent taxa Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum
Sibling taxa
A. aridum, A. arnowiae, A. contractum, A. coronatum, A. curvifolium, A. diegoense, A. eminens, A. hendersonii, A. hymenoides, A. latiglume, A. lemmonii, A. lettermanii, A. lobatum, A. nelsonii, A. nevadense, A. parishii, A. perplexum, A. pinetorum, A. richardsonii, A. robustum, A. scribneri, A. splendens, A. stillmanii, A. swallenii, A. thurberianum, A. wallowaense, A. webberi, A. ×bloomeri
A. aridum, A. contractum, A. coronatum, A. curvifolium, A. diegoense, A. eminens, A. hendersonii, A. hymenoides, A. latiglume, A. lemmonii, A. lettermanii, A. lobatum, A. nelsonii, A. nevadense, A. occidentale, A. parishii, A. perplexum, A. pinetorum, A. richardsonii, A. robustum, A. scribneri, A. splendens, A. stillmanii, A. swallenii, A. thurberianum, A. wallowaense, A. webberi, A. ×bloomeri
Subordinate taxa
A. occidentale subsp. californicum, A. occidentale subsp. occidentale, A. occidentale subsp. pubescens
Synonyms Stipa occidentalis, Stipa occidentalis var. montana
Name authority (Thurb.) Barkworth (S.L. Welsh & N.D Atwood) Barkworth
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