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Columbia needle grass, Nelson achnatherum, Nelson's needlegrass

Southwestern needlegrass

Habit Plants cespitose, not rhizomatous. Plants cespitose, shortly rhizomatous, bases knotty.
Culms

40-175 cm tall, 0.7-2.4 mm thick, lower cauline internodes usually glabrous, sometimes slightly pubescent below the lower nodes;

nodes 2-5.

50-100 cm tall, 0.8-1.5 mm thick, glabrous;

nodes 2-3.

Panicles

9-36 cm long, 0.8-2 cm wide;

branches ascending to appressed, straight.

20-55 cm long, 3-8 cm wide, open, often enclosed to midlength at anthesis;

lower branches 5-8 cm, ascending to divergent, flexuous.

Spikelets

appressed to the branches.

Glumes

6-12.5 mm long, 0.7-1.1 mm wide;

lower glumes exceeding the upper glumes by 0.2-0.8 mm;

florets 4.5-7 mm long, 0.6-0.9 mm thick, fusiform;

calluses 0.2-1 mm, blunt to sharp, dorsal boundary of the glabrous tip with the callus hairs almost straight to acute;

lemmas evenly hairy, hairs at midlength 0.5-1 mm, hairs at the apices to 2 mm, erect to ascending, apical lobes 0.1-0.4 mm, membranous, flexible;

awns 19-45 mm, persistent, twice-geniculate, first 2 segments scabrous or with hairs shorter than 0.5 mm, terminal segment straight;

paleas 2-4 mm, 1/3 – 2/3 as long as the lemmas, pubescent, hairs usually not exceeding the apices, veins terminating before the apices, apices rounded;

anthers 2-3.5 mm, dehiscent, not penicillate.

Lower glumes

5-12 mm long, 0.5-0.7 mm wide, 3-5-veined;

upper glumes 1-4 mm shorter, 3-veined;

florets 4-7.5 mm long, 0.5-0.9 mm thick, fusiform, terete;

calluses 1-2 mm, sharp;

lemmas evenly hairy, hairs 0.4-0.8 mm throughout, apical lobes not present;

awns 35-70 mm, persistent, twice-geniculate, first 2 segments scabrous, terminal segment flexuous;

paleas 1-2 mm, 1/3 – 1/2 as long as the lemmas, sparsely to moderately pubescent, apices rounded, flat;

anthers 3-3.5 mm, dehiscent, a few penicillate, hairs about 0.3 mm.

Caryopses

3-4 mm, fusiform.

about 4 mm, fusiform.

Basal

sheaths glabrous or sparsely to densely pubescent, margins sometimes ciliate;

collars glabrous or somewhat pubescent, without tufts of hair on the sides, collars of the flag leaves glabrous or sparsely pubescent;

basal ligules 0.2-0.7 mm, membranous, truncate to rounded, usually not ciliate;

upper ligules 1-1.5 mm, acute;

blades (0.5)1.2-5 mm wide.

sheaths mostly glabrous, ciliate on the margins;

collars glabrous on the back, usually with tufts of hair on the sides, hairs about 0.8 mm;

basal ligules 0.8-1.6 mm, membranous, glabrous, rounded to acute;

upper ligules to 4.5 mm, acute;

blades 0.7-3.5 mm wide, abaxial surfaces smooth to scabridulous, adaxial surfaces prominently ribbed, scabridulous or sparsely to densely pubescent, hairs about 0.1 mm.

2n

= 36, 44.

= 44, 46.

Achnatherum nelsonii

Achnatherum eminens

Distribution
from FNA
AZ; CA; CO; ID; MT; NM; NV; OR; SD; TX; UT; WA; WY; AB; BC; SK; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM; TX
[WildflowerSearch map]
[BONAP county map]
Discussion

Achnatherum nelsonii grows in meadows and openings, from sagebrush steppe and pinyon-juniper woodlands to subalpine forests, at 500-3500 m. It flowers in late spring to early summer, differing in this respect from A. perplexum. It is sometimes sympatric with A. lettermanii, from which it differs in its shorter paleas and wider leaves, and its tendency to grow in deeper or less disturbed soils. It differs from A. lemmonii in having wider leaf blades, shorter paleas, and membranous lemma lobes, and from A. nevadense and A. occidentale in its scabrous awns and the truncate to acute boundary of the glabrous tip of the callus with the callus hairs.

The two subspecies intergrade to some extent. There is also intergradation with Achnatherum occidentale, possibly as a result of hybridization and introgression.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Achnatherum eminens grows on dry, rocky slopes and valleys in the mountains of the southwestern United States, primarily in desert scrub, at 600-2600 m. Its range extends into Mexico. It is easy to recognize because of its open panicle, flexuous branches, and flexuous awns. It is superficially similar to Nassella cernua, but differs in its longer, glabrous ligules, not or weakly overlapping lemma margins, pubescent paleas, and geographic distribution.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Calluses blunt, dorsal boundary of the glabrous tip and the callus hairs almost straight to rounded; awns 19-31 mm long
subsp. dorei
1. Calluses sharp, dorsal boundary of the glabrous tip and the callus hairs acute; awns 19-45 mm long
subsp. nelsonii
Source FNA vol. 24, p. 123. FNA vol. 24, p. 133.
Parent taxa Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum
Sibling taxa
A. aridum, A. arnowiae, A. contractum, A. coronatum, A. curvifolium, A. diegoense, A. eminens, A. hendersonii, A. hymenoides, A. latiglume, A. lemmonii, A. lettermanii, A. lobatum, A. nevadense, A. occidentale, A. parishii, A. perplexum, A. pinetorum, A. richardsonii, A. robustum, A. scribneri, A. splendens, A. stillmanii, A. swallenii, A. thurberianum, A. wallowaense, A. webberi, A. ×bloomeri
A. aridum, A. arnowiae, A. contractum, A. coronatum, A. curvifolium, A. diegoense, A. hendersonii, A. hymenoides, A. latiglume, A. lemmonii, A. lettermanii, A. lobatum, A. nelsonii, A. nevadense, A. occidentale, A. parishii, A. perplexum, A. pinetorum, A. richardsonii, A. robustum, A. scribneri, A. splendens, A. stillmanii, A. swallenii, A. thurberianum, A. wallowaense, A. webberi, A. ×bloomeri
Subordinate taxa
A. nelsonii subsp. dorei, A. nelsonii subsp. nelsonii
Synonyms Stipa williamsii, Stipa occidentalis var. nelsonii, Stipa nelsonii Stipa eminens
Name authority (Scribn.) Barkworth (Cav.) Barkworth
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