Achnatherum nelsonii |
Achnatherum coronatum |
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Columbia needle grass, Nelson achnatherum, Nelson's needlegrass |
crested needlegrass, giant ricegrass |
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Habit | Plants cespitose, not rhizomatous. | Plants loosely cespitose, shortly rhizomatous, bases knotty. | ||||
Culms | 40-175 cm tall, 0.7-2.4 mm thick, lower cauline internodes usually glabrous, sometimes slightly pubescent below the lower nodes; nodes 2-5. |
55-210 cm tall, 3-6 mm thick, internodes usually glabrous, lower internodes sometimes puberulent; nodes 1-2, glabrous. |
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Panicles | 9-36 cm long, 0.8-2 cm wide; branches ascending to appressed, straight. |
15-60 cm long, 2-4 cm wide; branches widely spreading to ascending, longest branches 4-13 cm. |
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Spikelets | appressed to the branches. |
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Glumes | 6-12.5 mm long, 0.7-1.1 mm wide; lower glumes exceeding the upper glumes by 0.2-0.8 mm; florets 4.5-7 mm long, 0.6-0.9 mm thick, fusiform; calluses 0.2-1 mm, blunt to sharp, dorsal boundary of the glabrous tip with the callus hairs almost straight to acute; lemmas evenly hairy, hairs at midlength 0.5-1 mm, hairs at the apices to 2 mm, erect to ascending, apical lobes 0.1-0.4 mm, membranous, flexible; awns 19-45 mm, persistent, twice-geniculate, first 2 segments scabrous or with hairs shorter than 0.5 mm, terminal segment straight; paleas 2-4 mm, 1/3 – 2/3 as long as the lemmas, pubescent, hairs usually not exceeding the apices, veins terminating before the apices, apices rounded; anthers 2-3.5 mm, dehiscent, not penicillate. |
lanceolate, glabrous, tapering to awnlike apices; lower glumes 16-21 mm long, 1-1.3 mm wide, midveins scabrous; upper glumes 11-18 mm; florets 6.5-10 mm long, about 1 mm thick, fusiform, terete; calluses 0.5-2 mm, blunt to acute; lemmas densely hairy, hairs at midlength 1.5-4 mm, apical hairs 2-5 mm; awns 25-45 mm, persistent, twice-geniculate, all segments scabrous, terminal segment straight; paleas 3.5-5.5 mm, 3/5 – 9/10 as long as the lemmas, sparsely hairy between the veins, apices flat, rounded; anthers 3-4 mm, dehiscent, not penicillate. |
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Caryopses | 3-4 mm, fusiform. |
5-7 mm, fusiform. |
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Basal | sheaths glabrous or sparsely to densely pubescent, margins sometimes ciliate; collars glabrous or somewhat pubescent, without tufts of hair on the sides, collars of the flag leaves glabrous or sparsely pubescent; basal ligules 0.2-0.7 mm, membranous, truncate to rounded, usually not ciliate; upper ligules 1-1.5 mm, acute; blades (0.5)1.2-5 mm wide. |
sheaths mostly glabrous, often puberulent on the lower portion, flat, ribbonlike with age, margins hairy distally, hairs 1-2.5 mm; collars mostly glabrous; ligules 0.4-1.6(3) mm, truncate to slightly rounded, abaxial surfaces pubescent, ciliate, cilia about 0.5 mm; blades usually flat, 2.5-7 mm wide, both surfaces scabrous. |
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2n | = 36, 44. |
= 40. |
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Achnatherum nelsonii |
Achnatherum coronatum |
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Distribution |
AZ; CA; CO; ID; MT; NM; NV; OR; SD; TX; UT; WA; WY; AB; BC; SK; YT
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CA
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Discussion | Achnatherum nelsonii grows in meadows and openings, from sagebrush steppe and pinyon-juniper woodlands to subalpine forests, at 500-3500 m. It flowers in late spring to early summer, differing in this respect from A. perplexum. It is sometimes sympatric with A. lettermanii, from which it differs in its shorter paleas and wider leaves, and its tendency to grow in deeper or less disturbed soils. It differs from A. lemmonii in having wider leaf blades, shorter paleas, and membranous lemma lobes, and from A. nevadense and A. occidentale in its scabrous awns and the truncate to acute boundary of the glabrous tip of the callus with the callus hairs. The two subspecies intergrade to some extent. There is also intergradation with Achnatherum occidentale, possibly as a result of hybridization and introgression. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Achnatherum coronatum grows on gravel and on rocky slopes, mostly in chaparral associations of the Coast Range from Monterey County, California, to Baja California, Mexico. It is similar in size to A. diegoense, but differs in its mostly glabrous internodes and longer paleas. It differs from A. parishii, an inland species, in its twice-geniculate awns, more robust habit, and more sparsely pubescent paleas. Occasional plants combine the characteristics of both species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24, p. 123. | FNA vol. 24, p. 127. | ||||
Parent taxa | Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum | Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Stipa williamsii, Stipa occidentalis var. nelsonii, Stipa nelsonii | Stipa coronata | ||||
Name authority | (Scribn.) Barkworth | (Thurb.) Barkworth | ||||
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