Achnatherum diegoense |
Achnatherum richardsonii |
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San Diego County needlegrass, San Diego needlegrass |
Richardson needlegrass, Richardson's needlegrass, Richardson's rice grass, spreading needlegrass |
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Habit | Plants cespitose, not rhizomatous. | Plants tightly cespitose, not rhizomatous. |
Culms | 110-140 cm tall, 2.5-4 mm thick, internodes densely and retrorsely pubescent for 3-9 mm below the nodes, particularly the lower nodes, glabrous or retrorsely puberulent elsewhere; nodes 3, pubescent or glabrate. |
30-100 cm tall, 1-1.5 mm thick, glabrous; nodes usually 3. |
Panicles | 21-25 cm long, (2)4-8 cm wide; branches strongly divergent to ascending, straight, lower branches 5-7 cm. |
7-25 cm long, 7-15 cm wide; branches divergent, flexuous, longest branches 7-10 cm, with the spikelets confined to the distal 1/4. |
Spikelets | appressed to the branches. |
pendulous. |
Glumes | subequal, 8-11.5 mm; lower glumes 0.5-1 mm wide, 3-5-veined; upper glumes 3-veined; florets 5.5-7.5 mm long, 0.7-1 mm thick, fusiform, terete; calluses 0.25-1.2 mm, acute; lemmas evenly hairy, hairs at midlength and at the apices 0.5-1 mm, apical lobes 0.2-0.4 mm, membranous, flexible; awns 20-50 mm, persistent, twice-geniculate, all segments scabrous to scabridulous, terminal segment straight; paleas 2.6-3.8 mm, 1/2 - 3/4 as long as the lemmas, pubescent, hairs not extending beyond the apices, veins terminating below the apices, apices rounded; anthers 2.5-4 mm, dehiscent, not penicillate. |
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Lower glumes | 7.5-11 mm long, 0.9-1.2 mm wide; upper glumes 2-3 mm shorter; florets 5-6 mm long, 0.6-0.9 mm thick, fusiform, terete; calluses 0.4-0.7 mm, blunt; lemmas evenly hairy on the lower portion, often glabrate distally, body and apical hairs 0.2-0.5 mm, apical lobes not or scarcely developed, to 0.1 mm; awns 15-25 mm, persistent, twice-geniculate, first 2 segments strigulose, hairs about 0.1 mm, terminal segment straight; paleas 2.2-3.6 mm, 1/2 - 3/5 as long as the lemmas, pubescent, hairs not exceeding the apices, apices rounded; anthers 2.5-3 mm, dehiscent, penicillate, hairs 0.1-0.5 mm. |
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Caryopses | 3-4 mm, fusiform. |
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Basal | sheaths mostly glabrous or puberulent, margins ciliate distally; collars glabrous or with hairs, hairs mostly to 0.5 mm, sides with tufts of 1.5-2 mm hairs; ligules 0.4-2 mm, rounded to acute, abaxial surfaces hairy, hairs to 0.5 mm; upper ligules 1-3 mm, similar in structure and pubescence; blades 1-3.5 mm wide, abaxial surfaces smooth or scabrous, adaxial surfaces prominently ribbed, hairy, hairs 2-3 mm. |
sheaths glabrous, margins ciliolate; collars glabrous, without tufts of hair on the sides; ligules 0.1-0.5 mm, truncate, ciliolate; blades 0.8-3 mm wide, convolute when dry, abaxial surfaces scabridulous, adaxial surfaces glabrous. |
2n | = unknown. |
= 44. |
Achnatherum diegoense |
Achnatherum richardsonii |
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Distribution |
CA
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AK; CO; ID; MT; SD; WA; WY; AB; BC; MB; SK; YT
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Discussion | Achnatherum diegoense grows in chaparral and coastal sage scrub, on rocky soil near streams or the coast, at 0-350 m, on the Channel Islands of Santa Barbara County, California, and, on the mainland, in Ventura and San Diego counties south into Baja California, Mexico. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Achnatherum richardsonii grows in open woodlands and grasslands,often on sand or gravel, from the Yukon Territory to Washington and Manitoba, and south in the Rocky Mountains through Montana and Wyoming to western South Dakota and northern Colorado. Its elevation range is 1000-3100 m. It is readily recognized by its combination of flexuous panicle branches, drooping spikelets, and straight distal awn segments. Scagel and Maze (1984) concluded that putative hybrids between A. richardsonii and A. nelsonii subsp. dorei were merely large plants of subsp. dorei that varied in the direction of A. richardsonii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 24, p. 131. | FNA vol. 24, p. 133. |
Parent taxa | Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum | Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum |
Sibling taxa | ||
Synonyms | Stipa richardsonii | |
Name authority | (Swallen) Barkworth | (Link) Barkworth |
Web links |
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