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San Diego County needlegrass, San Diego needlegrass

Nevada needlegrass

Habit Plants cespitose, not rhizomatous. Plants cespitose, not rhizomatous.
Culms

110-140 cm tall, 2.5-4 mm thick, internodes densely and retrorsely pubescent for 3-9 mm below the nodes, particularly the lower nodes, glabrous or retrorsely puberulent elsewhere;

nodes 3, pubescent or glabrate.

20-85 cm tall, 0.8-1.2 mm thick, usually retrorsely pubescent below the lower nodes, sometimes glabrous, sometimes pubescent over the whole of the internodes;

nodes 3-4.

Panicles

21-25 cm long, (2)4-8 cm wide;

branches strongly divergent to ascending, straight, lower branches 5-7 cm.

6-25 cm long, 0.4-1.5 cm wide;

branches appressed, lower branches 2-7.5 cm.

Spikelets

appressed to the branches.

appressed to the branches.

Glumes

subequal, 8-11.5 mm;

lower glumes 0.5-1 mm wide, 3-5-veined;

upper glumes 3-veined;

florets 5.5-7.5 mm long, 0.7-1 mm thick, fusiform, terete;

calluses 0.25-1.2 mm, acute;

lemmas evenly hairy, hairs at midlength and at the apices 0.5-1 mm, apical lobes 0.2-0.4 mm, membranous, flexible;

awns 20-50 mm, persistent, twice-geniculate, all segments scabrous to scabridulous, terminal segment straight;

paleas 2.6-3.8 mm, 1/2 - 3/4 as long as the lemmas, pubescent, hairs not extending beyond the apices, veins terminating below the apices, apices rounded;

anthers 2.5-4 mm, dehiscent, not penicillate.

subequal, 7-14 mm;

lower glumes 0.6-0.9 mm wide;

florets 5-6.5 mm long, 0.6-1 mm thick, fusiform, terete;

calluses 0.5-0.7 mm, sharp, dorsal boundary of the glabrous tip with the callus hairs rounded to acute;

lemmas evenly hairy, hairs 0.5-1 mm at midlength, apical hairs 0.8-2 mm, longer than those at midlength and those at the base of the awn, apical lobes membranous, about 0.2 mm;

awns 20-35 mm, persistent, twice-geniculate, first 2 segments pilose, with hairs 0.5-1.5 mm and of mixed lengths, terminal segment scabridulous to smooth;

paleas 2.8-4.2 mm, 1/2 - 3/4 as long as the lemmas, pubescent, distal hairs usually about 1 mm, extending well beyond the apices, apices rounded;

anthers about 2.5 mm, dehiscent, not penicillate.

Caryopses

3-5.5 mm, fusiform.

Basal

sheaths mostly glabrous or puberulent, margins ciliate distally;

collars glabrous or with hairs, hairs mostly to 0.5 mm, sides with tufts of 1.5-2 mm hairs;

ligules 0.4-2 mm, rounded to acute, abaxial surfaces hairy, hairs to 0.5 mm;

upper ligules 1-3 mm, similar in structure and pubescence;

blades 1-3.5 mm wide, abaxial surfaces smooth or scabrous, adaxial surfaces prominently ribbed, hairy, hairs 2-3 mm.

sheaths glabrous or pubescent, sometimes scabridulous, usually glabrous at the throat, becoming brown to gray-brown;

collars, including the sides, glabrous;

basal ligules 0.2-0.7 mm, truncate;

upper ligules 0.3-1 mm, often wider than the blades;

blades usually 10-25 cm long, 1-3 mm wide, usually involute, abaxial surfaces glabrous, adaxial surfaces more or less puberulent.

2n

= unknown.

= 68.

Achnatherum diegoense

Achnatherum nevadense

Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CO; ID; NV; OR; UT; WY
[BONAP county map]
Discussion

Achnatherum diegoense grows in chaparral and coastal sage scrub, on rocky soil near streams or the coast, at 0-350 m, on the Channel Islands of Santa Barbara County, California, and, on the mainland, in Ventura and San Diego counties south into Baja California, Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Achnatherum nevadense grows in sagebrush and open woodlands, from Washington to south-central Wyoming and south to California and Utah. Johnson (1962) argued that it is an alloploid derivative of A. occidentale and A. lettermanii.

The apical lemma hairs of Achnatherum nevadense appear longer than the lowermost awn hairs. This difference is not reflected in the lengths shown because a few of the basal awn hairs may be as long as those at the top of the lemma, but the majority are shorter. This is the best character for distinguishing A. nevadense and A. occidentale subsp. californicum from A. occidentale subsp. pubescens. In addition, in A. nevadense and A. occidentale subsp. californicum, the hairs on the first awn segments tend to look untidy because of their varied lengths and the different angles they make with the awn; those of A. occidentale subsp. occidentale and subsp. pubescens have basal awn segments with tidier looking hairs.

Differentiating between Achnatherum nevadense and A. occidentale subsp. californicutn can be difficult, but they differ in the shape of the boundary between the glabrous and strigose portions of the callus. In addition, A. nevadense is usually pubescent below the lower cauline nodes, and has paleas that are longer in relation to the lemmas. Plants of A. nevadense from Convict Creek Basin, California, are unusual in having culms that are glabrous below the lower nodes, but in other respects they correspond to A. nevadense. Achnatherum nevadense also resembles A. latiglume, but the latter species has blunter calluses and paleas that tend to be thicker and somewhat longer in comparison to the lemmas than those of A. nevadense.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 131. FNA vol. 24, p. 120.
Parent taxa Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum
Sibling taxa
A. aridum, A. arnowiae, A. contractum, A. coronatum, A. curvifolium, A. eminens, A. hendersonii, A. hymenoides, A. latiglume, A. lemmonii, A. lettermanii, A. lobatum, A. nelsonii, A. nevadense, A. occidentale, A. parishii, A. perplexum, A. pinetorum, A. richardsonii, A. robustum, A. scribneri, A. splendens, A. stillmanii, A. swallenii, A. thurberianum, A. wallowaense, A. webberi, A. ×bloomeri
A. aridum, A. arnowiae, A. contractum, A. coronatum, A. curvifolium, A. diegoense, A. eminens, A. hendersonii, A. hymenoides, A. latiglume, A. lemmonii, A. lettermanii, A. lobatum, A. nelsonii, A. occidentale, A. parishii, A. perplexum, A. pinetorum, A. richardsonii, A. robustum, A. scribneri, A. splendens, A. stillmanii, A. swallenii, A. thurberianum, A. wallowaense, A. webberi, A. ×bloomeri
Synonyms Stipa nevadetisis
Name authority (Swallen) Barkworth (B.L. Johnson) Barkworth
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