Achnatherum diegoense |
Achnatherum nelsonii |
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San Diego County needlegrass, San Diego needlegrass |
Columbia needle grass, Nelson achnatherum, Nelson's needlegrass |
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Habit | Plants cespitose, not rhizomatous. | Plants cespitose, not rhizomatous. | ||||
Culms | 110-140 cm tall, 2.5-4 mm thick, internodes densely and retrorsely pubescent for 3-9 mm below the nodes, particularly the lower nodes, glabrous or retrorsely puberulent elsewhere; nodes 3, pubescent or glabrate. |
40-175 cm tall, 0.7-2.4 mm thick, lower cauline internodes usually glabrous, sometimes slightly pubescent below the lower nodes; nodes 2-5. |
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Panicles | 21-25 cm long, (2)4-8 cm wide; branches strongly divergent to ascending, straight, lower branches 5-7 cm. |
9-36 cm long, 0.8-2 cm wide; branches ascending to appressed, straight. |
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Spikelets | appressed to the branches. |
appressed to the branches. |
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Glumes | subequal, 8-11.5 mm; lower glumes 0.5-1 mm wide, 3-5-veined; upper glumes 3-veined; florets 5.5-7.5 mm long, 0.7-1 mm thick, fusiform, terete; calluses 0.25-1.2 mm, acute; lemmas evenly hairy, hairs at midlength and at the apices 0.5-1 mm, apical lobes 0.2-0.4 mm, membranous, flexible; awns 20-50 mm, persistent, twice-geniculate, all segments scabrous to scabridulous, terminal segment straight; paleas 2.6-3.8 mm, 1/2 - 3/4 as long as the lemmas, pubescent, hairs not extending beyond the apices, veins terminating below the apices, apices rounded; anthers 2.5-4 mm, dehiscent, not penicillate. |
6-12.5 mm long, 0.7-1.1 mm wide; lower glumes exceeding the upper glumes by 0.2-0.8 mm; florets 4.5-7 mm long, 0.6-0.9 mm thick, fusiform; calluses 0.2-1 mm, blunt to sharp, dorsal boundary of the glabrous tip with the callus hairs almost straight to acute; lemmas evenly hairy, hairs at midlength 0.5-1 mm, hairs at the apices to 2 mm, erect to ascending, apical lobes 0.1-0.4 mm, membranous, flexible; awns 19-45 mm, persistent, twice-geniculate, first 2 segments scabrous or with hairs shorter than 0.5 mm, terminal segment straight; paleas 2-4 mm, 1/3 – 2/3 as long as the lemmas, pubescent, hairs usually not exceeding the apices, veins terminating before the apices, apices rounded; anthers 2-3.5 mm, dehiscent, not penicillate. |
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Caryopses | 3-4 mm, fusiform. |
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Basal | sheaths mostly glabrous or puberulent, margins ciliate distally; collars glabrous or with hairs, hairs mostly to 0.5 mm, sides with tufts of 1.5-2 mm hairs; ligules 0.4-2 mm, rounded to acute, abaxial surfaces hairy, hairs to 0.5 mm; upper ligules 1-3 mm, similar in structure and pubescence; blades 1-3.5 mm wide, abaxial surfaces smooth or scabrous, adaxial surfaces prominently ribbed, hairy, hairs 2-3 mm. |
sheaths glabrous or sparsely to densely pubescent, margins sometimes ciliate; collars glabrous or somewhat pubescent, without tufts of hair on the sides, collars of the flag leaves glabrous or sparsely pubescent; basal ligules 0.2-0.7 mm, membranous, truncate to rounded, usually not ciliate; upper ligules 1-1.5 mm, acute; blades (0.5)1.2-5 mm wide. |
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2n | = unknown. |
= 36, 44. |
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Achnatherum diegoense |
Achnatherum nelsonii |
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Distribution |
CA
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AZ; CA; CO; ID; MT; NM; NV; OR; SD; TX; UT; WA; WY; AB; BC; SK; YT
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Discussion | Achnatherum diegoense grows in chaparral and coastal sage scrub, on rocky soil near streams or the coast, at 0-350 m, on the Channel Islands of Santa Barbara County, California, and, on the mainland, in Ventura and San Diego counties south into Baja California, Mexico. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Achnatherum nelsonii grows in meadows and openings, from sagebrush steppe and pinyon-juniper woodlands to subalpine forests, at 500-3500 m. It flowers in late spring to early summer, differing in this respect from A. perplexum. It is sometimes sympatric with A. lettermanii, from which it differs in its shorter paleas and wider leaves, and its tendency to grow in deeper or less disturbed soils. It differs from A. lemmonii in having wider leaf blades, shorter paleas, and membranous lemma lobes, and from A. nevadense and A. occidentale in its scabrous awns and the truncate to acute boundary of the glabrous tip of the callus with the callus hairs. The two subspecies intergrade to some extent. There is also intergradation with Achnatherum occidentale, possibly as a result of hybridization and introgression. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24, p. 131. | FNA vol. 24, p. 123. | ||||
Parent taxa | Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum | Poaceae > subfam. Pooideae > tribe Stipeae > Achnatherum | ||||
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Subordinate taxa | ||||||
Synonyms | Stipa williamsii, Stipa occidentalis var. nelsonii, Stipa nelsonii | |||||
Name authority | (Swallen) Barkworth | (Scribn.) Barkworth | ||||
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