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Paraguay starburr, Paraguayan starbur, Paraguayan starburr, southern starbur

starbur, starburr

Habit Plants 10–60(–120+) cm. Annuals (sometimes persisting), 10–60(–120) cm.
Stems

± procumbent.

erect to ± prostrate (repeatedly “forked”).

Leaves

blades deltate to ± rhombic or ovate, 13–37 × 7–32 mm, faces sparsely scabrellous to glabrate or glabrescent, gland-dotted.

cauline; opposite; petiolate or ± sessile;

blades mostly elliptic to deltate, rhombic, or ovate, sometimes lyrate, ultimate margins entire or toothed, faces usually pilosulous to sericeous or scabrellous, sometimes glabrate or glabrescent, usually gland-dotted.

Involucres

± hemispheric, 3–5 mm diam. (becoming ± rotate in fruit).

Receptacles

convex, paleate (paleae cuneate to spatulate, ± conduplicate or flattish, membranous).

Ray florets

5–8, pistillate, fertile;

corollas yellowish (tubes shorter than to equaling laminae, laminae ovate to elliptic or linear).

Disc florets

3–8(–12+), functionally staminate;

corollas yellowish, tubes shorter than funnelform or campanulate throats, lobes 5, deltate.

Phyllaries

persistent (outer) or falling, 10–13 in 2 series (outer 4–6 herbaceous, inner 5–8 each investing a ray ovary, enlarging in fruit to form a perigynium, shed with enclosed cypsela).

Fruits

plumply ellipsoid to fusiform, weakly compressed, 7–9+ mm, 5–7-ribbed, lacking terminal spines, prickles ± uncinate, mostly along ribs.

Heads

radiate, 1(–3) in “forks” of branches (terminal, appearing axillary by sympodial growth).

Cypselae

each enclosed within and shed with an often hardened, ± prickly perigynium (the ultimate “fruits” plumply ellipsoid to fusiform, or ± compressed);

pappi 0 or rudimentary.

x

= 11.

2n

= 22.

Acanthospermum australe

Acanthospermum

Phenology Flowering year round, mostly Jul–Aug.
Habitat Disturbed, often sandy sites
Elevation 0–300 m (0–1000 ft)
Distribution
from FNA
AL; AR; DC; FL; GA; LA; MA; MS; NC; OR; PA; SC; TX; VA; South America [Introduced in North America; also introduced in Mexico, West Indies, Central America, Europe, Asia, Africa, Pacific Islands (Hawaii)]
[WildflowerSearch map]
[BONAP county map]
from USDA
mostly tropical to warm-temperate New World [Introduced in North America; also introduced in Old World]
[BONAP county map]
Discussion

Reports of Acanthospermum xanthioides (Kunth) de Candolle from New York are evidently based on specimens of A. australe.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 6 (3 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruits 7–9+ mm, 5–7-ribbed, terminal spines 0
A. australe
1. Fruits 2–6 mm, 3-ribbed or not notably ribbed, terminal spines 2
→ 2
2. Leaf blades rhombic-ovate to obovate, (20–)40–120(–150+) mm; fruits not notably ribbed, prickles ± scattered
A. hispidum
2. Leaf blades ovate to lyrate, 10–30(–45) mm; fruits usually 3-ribbed, prickles mostly along 2 ribs and around apices
A. humile
Source FNA vol. 21, p. 37. FNA vol. 21, p. 36. Author: John L. Strother.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Melampodiinae > Acanthospermum Asteraceae > tribe Heliantheae > subtribe Melampodiinae
Sibling taxa
A. hispidum, A. humile
Subordinate taxa
A. australe, A. hispidum, A. humile
Synonyms Melampodium australe
Name authority (Loefling) Kuntze: Revis. Gen. Pl. 1: 303. (1891) Schrank: Pl. Rar. Hort. Monac. 2: plate 53. (1820)
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