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flowery puncturebract

flowery puncturebract, Parish's oxytheca

Habit Herbs, annual; taproot slender. Plants (0.5–)1–6 × 0.5–4 dm.
Stems

arising directly from the root, erect to somewhat spreading, solid, not fistulose or disarticulating into ringlike segments, glabrous, partially glandular.

Leaves

usually persistent through anthesis, basal, rosulate;

petiole indistinct;

blade broadly obovate or spatulate to oblong or oblanceolate, margins ciliate-denticulate.

blades 1–7 × 0.5–2 cm.

Inflorescences

terminal, cymose;

branches dichotomous, not brittle or disarticulating into segments, round, glabrous except for glandular nodes and proximal 1/2 of internode;

bracts 2–3, connate basally, scalelike, triangular to lanceolate or subulate, awned at proximal nodes, glabrous or sparsely glandular.

1–5 dm;

bracts 1–4(–10) × 0.5–3 mm, awns 0.2–1.5 mm.

Peduncles

erect to deflexed.

(0.3–)1–5(–7.5) cm.

Involucres

1 per node, not ribbed, tubular, turbinate;

teeth 4(–5) or 7–30(–36), 1/2 to completely connate, awn-tipped, glandular distally adaxially.

1.5–2 mm, awns ivory or reddish, 2–5 mm.

Flowers

3–10(–20) per involucre at any single time during full anthesis;

perianth white to rose, occasionally greenish white to cream, broadly campanulate when open, narrowly urceolate when closed, hirsute to strigose and sparsely glandular abaxially;

tepals 6, connate 1/4–1/3 their length, monomorphic, entire or rarely irregularly divided apically;

stamens 9;

filaments basally adnate, glabrous or sparsely pubescent basally;

anthers red to maroon, oblong to oval.

perianth 2–2.5 mm;

filaments 2–2.5 mm;

anthers 0.3–0.6 mm.

Achenes

usually included, dark brown to deep maroon, not winged, globose-lenticular, glabrous.

1.7–2 mm.

Seeds

embryo curved.

x

= 20.

Acanthoscyphus

Acanthoscyphus parishii

Distribution
s Calif
[BONAP county map]
s Calif
[WildflowerSearch map]
[BONAP county map]
Discussion

Species 1.

Acanthoscyphus is allied to Eriogonum subg. Ganysma, specifically to E. parishii, E. apiculatum, and their close relatives. It is distinguished from Eriogonum by its awned involucres.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 4 (4 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Involucral awns 7-16, dark red, 3-4 mm; San Rafael Mountains, Topatopa Mountain, and Mount Pinos, Santa Barbara and Ventura counties, California
var. abramsii
1. Involucral awns 4-36, ivory colored, 2-5 mm; San Bernardino and Ventura counties
→ 2
2. Involucral awns 4(-5), 2-3 mm; ne side of San Bernardino Mountains, San Bernardino County, California
var. goodmaniana
2. Involucral awns 7-30(-36), 3-5 mm; se side of e San Bernardino County, and Ventura County, California
→ 3
3. Involucral awns (10-)13-30(-36); inflorescence bracts triangular, awns 0.2-0.5 mm; Pine Mountain, Ventura County, e to c San Bernardino Mountains, San Bernardino County, California
var. parishii
3. Involucral awns 7-10; inflorescence bracts subulate, awns 0.5-1.5 mm; se side of e San Bernardino Mountains, San Bernardino County, California
var. cienegensis
Source FNA vol. 5, p. 437. Author: James L. Reveal. FNA vol. 5, p. 438.
Parent taxa Polygonaceae > subfam. Eriogonoideae Polygonaceae > subfam. Eriogonoideae > Acanthoscyphus
Subordinate taxa
A. parishii
A. parishii var. abramsii, A. parishii var. cienegensis, A. parishii var. goodmaniana, A. parishii var. parishii
Synonyms Oxytheca section A. Oxytheca parishii
Name authority Small: Bull. Torrey Bot. Club 25: 53. (1898) (Parry) Small: Bull. Torrey Bot. Club 25: 53. (1898)
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