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beefsteak plant, match-me-if-you-can, painted copperleaf, wilkes' acalypha

Virginia copperleaf, Virginia three-seed-Mercury

Habit Shrubs, 20–50 dm, monoecious. Herbs, annual, 1–5 dm, monoecious.
Stems

erect, sparsely to densely pubescent, not glandular.

usually erect, sometimes ascending, pubescent and usually hirsute.

Leaves

persistent;

petiole 1–6 cm;

blade ovate to broadly ovate or suborbiculate, 9–20 × 4–15 cm, base obtuse to rounded or subcordate, margins serrate-crenate, apex acuminate.

petiole 0.3–7 cm;

blade narrowly rhombic to broadly lanceolate, 1–8(–11) × 0.5–3(–4) cm, base acute, margins serrate, apex acute to acuminate.

Inflorescences

usually unisexual, rarely bisexual, axillary;

staminate peduncle 0.1–1.5 cm, fertile portion 10–20 cm;

pistillate peduncle 1–2 cm, fertile portion 4–15 × 0.5–0.8 cm;

bisexual similar to staminate, with 1–2 pistillate bracts near base;

allomorphic pistillate flowers absent.

bisexual, axillary;

peduncle 0.1–0.6 cm, pistillate portion 0.7–1.5 × 1.3–1.7 cm or pistillate bract solitary, staminate portion 0.3–1(–1.8) cm;

allomorphic pistillate flowers absent.

Pistillate flowers

pistil 3-carpellate;

styles multifid or laciniate.

pistil 3-carpellate;

styles multifid or laciniate.

Capsules

unknown.

smooth, pubescent.

Seeds

unknown.

(1.2–)1.5–1.6(–1.8) mm, minutely pitted.

Pistillate

bracts loosely arranged (inflorescence axis visible between bracts), 2–4 × 3–5 mm, abaxial surface sparsely to moderately pubescent;

lobes 7–9, ovate to lanceolate, 1/4 bract length, except terminal lobe to 1/2 bract length.

bracts loosely arranged to crowded (inflorescence axis clearly to scarcely visible between bracts) or solitary, 6–13 × 9–20 mm, abaxial surface hirsute and sometimes stipitate-glandular;

lobes (9–)10–14(–16), triangular, 1/4–1/2 bract length.

Acalypha wilkesiana

Acalypha virginica

Phenology Flowering spring–fall. Flowering and fruiting summer–fall.
Habitat Old home sites, disturbed areas. Deciduous and evergreen woods, riverbanks, agricultural fields, disturbed areas.
Elevation 0–10 m. (0–0 ft.) 30–1200 m. (100–3900 ft.)
Distribution
from FNA
FL; Pacific Islands [Introduced in North America; introduced also in Mexico, West Indies, Central America, n South America, se Asia, Africa]
from FNA
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; WV
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[BONAP county map]
Discussion

Acalypha wilkesiana is not known in the wild, but presumably originated in the southwestern Pacific Islands (Bismarck Archipelago east to Fiji). The species is commonly cultivated as an ornamental for its leaves that may be various shades of green, purple, red, orange, and yellow (sometimes variegated), and sometimes contorted into unusual shapes. Despite low seed set, it occasionally becomes naturalized in tropical and subtropical areas. Naturalized plants often lack the distinctive leaf coloration found in cultivated plants. Although sometimes treated as A. amentacea subsp. wilkesiana, DNA sequence data show that A. wilkesiana and A. amentacea are distinct species (V. G. Sagun et al. 2010).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Acalypha virginica overlaps geographically with A. rhomboidea but is more frequently found in grassy or prairielike habitats. See the discussion of 15. A. rhomboidea for notes on the nomenclatural and taxonomic confusion associated with these species.

Acalypha virginica and A. gracilens can sometimes be difficult to distinguish, especially when young. They can generally be distinguished by the pistillate bracts, which in A. virginica are hirsute and lack red sessile glands abaxially and in A. gracilens are sparsely pubescent and bear some red sessile glands abaxially.

Reports of Acalypha virginica from states other than those listed here are based on misidentifications.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 165. FNA vol. 12, p. 171.
Parent taxa Euphorbiaceae > Acalypha Euphorbiaceae > Acalypha
Sibling taxa
A. alopecuroidea, A. arvensis, A. australis, A. californica, A. chamaedrifolia, A. deamii, A. gracilens, A. monococca, A. monostachya, A. neomexicana, A. ostryifolia, A. phleoides, A. poiretii, A. radians, A. rhomboidea, A. setosa, A. virginica
A. alopecuroidea, A. arvensis, A. australis, A. californica, A. chamaedrifolia, A. deamii, A. gracilens, A. monococca, A. monostachya, A. neomexicana, A. ostryifolia, A. phleoides, A. poiretii, A. radians, A. rhomboidea, A. setosa, A. wilkesiana
Synonyms A. amentacea subsp. wilkesiana A. digyneia
Name authority Müller Arg.: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 15(2): 817. (1866) Linnaeus: Sp. Pl. 2: 1003. (1753) — name conserved
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