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hophornbeam copperleaf, pineland threeseed Mercury

California acalypha, California copperleaf, heirba del cancer, Pringle threeseed Mercury

Habit Herbs, annual, 3.5–7 dm, monoecious. Shrubs, 5–10 dm, monoecious.
Stems

erect, pubescent, sparsely hirsute, and stipitate-glandular.

erect, hirsute and stipitate-glandular, becoming glabrate.

Leaves

petiole 1.5–6.5 cm;

blade ovate, 3–8 × 1.5–5 cm, base cordate, margins serrate, apex acute to short acuminate.

persistent or drought-deciduous;

petiole 0.5–2.5 cm;

blade ovate to cordate, 1–5 × 0.5–4 cm, base truncate to rounded or cordate, margins serrate-crenate, apex acute or obtuse.

Inflorescences

unisexual, axillary (staminate) and terminal (pistillate; sometimes on short lateral branches, appearing axillary);

staminate peduncle 0.5–1.5 cm, fertile portion 0.5–3.5 cm;

pistillate peduncle 0.1–1 cm, fertile portion 3–7 × 0.7–1 cm;

allomorphic pistillate flowers common, near apices of pistillate inflorescences.

unisexual and bisexual, axillary and terminal;

staminate peduncle 0.3–2.5 cm, fertile portion 1–4 cm;

pistillate peduncle 0.4–3 cm, fertile portion 1–3 × 0.8–1.2 cm;

bisexual similar to staminate, with 1–3 pistillate bracts near base;

allomorphic pistillate flowers absent.

Pedicels

of allomorphic flowers rudimentary.

Pistillate flowers

pistil 3-carpellate (normal flowers), 1(–3)-carpellate (allomorphic flowers);

styles multifid or laciniate.

pistil 3-carpellate;

styles multifid or laciniate.

Capsules

spiny, pubescent;

allomorphic fruits obovoid, 2 irregular flanges near apex, 2–2.2 × 1.6–1.8 mm, smooth, pubescent.

smooth, pubescent and stipitate-glandular.

Seeds

1.6–2 mm, tuberculate.

1.5–2 mm, minutely pitted.

Pistillate

bracts (normal and allomorphic flowers) loosely arranged (inflorescence axis visible between bracts), 3–6 × 6–8 mm, abaxial surface pubescent (hairs to 0.3 mm) and sparsely stipitate-glandular;

lobes (9–)13–17, linear, 2/3 bract length, muricate.

bracts loosely arranged (inflorescence axis visible between bracts), 3–6 × 5.5–11 mm, abaxial surface pubescent, sessile- and stipitate-glandular;

lobes (8–)10–18, rounded, 1/5 bract length.

2n

= 20.

Acalypha ostryifolia

Acalypha californica

Phenology Flowering and fruiting summer–fall. Flowering and fruiting year-round, especially spring and fall.
Habitat Stream banks, edges of woods, disturbed areas, agricultural fields. Arid rocky slopes, desert washes.
Elevation 0–1700 m. (0–5600 ft.) 10–1400 m. (0–4600 ft.)
Distribution
from FNA
AL; AR; AZ; FL; GA; IA; IL; IN; KS; KY; LA; MO; MS; NC; NE; OH; OK; SC; TN; TX; VA; WV; Mexico; West Indies [Introduced to Central America]
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; Mexico (Baja California, Baja California Sur, Sinaloa, Sonora)
[WildflowerSearch map]
[BONAP county map]
Discussion

J. Torrey [in W. H. Emory 1857–1859, vol. 2(1)] reported Acalypha ostryifolia (as A. caroliniana Elliott) from New Mexico on the basis of Bigelow s.n., collected near "the Copper Mines" in Grant County. Although no specimen has been located by the author and this is the only report of this species from New Mexico, it is to be expected in the southwestern part of the state. It was collected in the late nineteenth century in New Jersey and Pennsylvania, but has not been collected there since.

Acalypha ostryifolia may not be native to the northern part of its range. Specimen collection dates suggest that the species is spreading northward, and in much of its range it is found primarily in areas with human disturbance.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Plants in Arizona and Sonora have been segregated as Acalypha pringlei based on having long nonglandular hairs mixed with shorter hairs on the stem (versus hairs all of one length). This trait appears throughout the range of A. californica and cannot be used to distinguish two species (G. A. Levin 1995).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 167. FNA vol. 12, p. 165.
Parent taxa Euphorbiaceae > Acalypha Euphorbiaceae > Acalypha
Sibling taxa
A. alopecuroidea, A. arvensis, A. australis, A. californica, A. chamaedrifolia, A. deamii, A. gracilens, A. monococca, A. monostachya, A. neomexicana, A. phleoides, A. poiretii, A. radians, A. rhomboidea, A. setosa, A. virginica, A. wilkesiana
A. alopecuroidea, A. arvensis, A. australis, A. chamaedrifolia, A. deamii, A. gracilens, A. monococca, A. monostachya, A. neomexicana, A. ostryifolia, A. phleoides, A. poiretii, A. radians, A. rhomboidea, A. setosa, A. virginica, A. wilkesiana
Synonyms A. pringlei
Name authority Riddell ex J. M. Coulter: Mem. Torrey Bot. Club 5: 213. (1894) — (as ostryaefolia) Bentham: Bot. Voy. Sulphur, 51. (1844)
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