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field copperleaf

hophornbeam copperleaf, pineland threeseed Mercury

Habit Herbs, annual, 2–8 dm, monoecious. Herbs, annual, 3.5–7 dm, monoecious.
Stems

erect to ascending, short-pubescent and densely [sparsely] hirsute.

erect, pubescent, sparsely hirsute, and stipitate-glandular.

Leaves

petiole 0.5–4 cm, not stipitate-glandular;

blade rhombic-ovate to rhombic-lanceolate, 2–9(–12) × 1.2–5(–6.5) cm, base cuneate to rounded, margins serrate, apex obtuse to acute.

petiole 1.5–6.5 cm;

blade ovate, 3–8 × 1.5–5 cm, base cordate, margins serrate, apex acute to short acuminate.

Inflorescences

unisexual or bisexual, axillary;

staminate peduncle 0.3–2.5 cm, not stipitate-glandular, fertile portion 1.5–3[–6] cm;

pistillate peduncle 0.4–3 cm, not stipitate-glandular, fertile portion 2.5–4[–8] × 1–2 cm;

bisexual similar to pistillate, with staminate portion 0.4–0.7 cm;

allomorphic pistillate flowers common, terminal on pistillate and bisexual inflorescences.

unisexual, axillary (staminate) and terminal (pistillate; sometimes on short lateral branches, appearing axillary);

staminate peduncle 0.5–1.5 cm, fertile portion 0.5–3.5 cm;

pistillate peduncle 0.1–1 cm, fertile portion 3–7 × 0.7–1 cm;

allomorphic pistillate flowers common, near apices of pistillate inflorescences.

Pedicels

of allomorphic flowers rudimentary if borne above staminate flowers or 10–18 mm if borne above normal pistillate flowers.

of allomorphic flowers rudimentary.

Pistillate flowers

pistil 3-carpellate (normal flowers), (1–)2-carpellate (allomorphic flowers);

styles multifid or laciniate.

pistil 3-carpellate (normal flowers), 1(–3)-carpellate (allomorphic flowers);

styles multifid or laciniate.

Capsules

smooth, hispidulous;

allomorphic fruits ovoid, 1.5–1.6 × 1.2–1.3 mm, muricate, sparsely to densely puberulent.

spiny, pubescent;

allomorphic fruits obovoid, 2 irregular flanges near apex, 2–2.2 × 1.6–1.8 mm, smooth, pubescent.

Seeds

1.1–1.5 mm, minutely pitted.

1.6–2 mm, tuberculate.

Pistillate

bracts (normal flowers) very densely crowded (inflorescence axis not visible between bracts), 6–12 × 4–6 mm, abaxial surface long-hirsute (hairs to 2 mm) and stipitate-glandular;

lobes 3–7, proximally deltate with linear tips, 1/2–2/3 bract length, smooth; of allomorphic flowers absent.

bracts (normal and allomorphic flowers) loosely arranged (inflorescence axis visible between bracts), 3–6 × 6–8 mm, abaxial surface pubescent (hairs to 0.3 mm) and sparsely stipitate-glandular;

lobes (9–)13–17, linear, 2/3 bract length, muricate.

Acalypha arvensis

Acalypha ostryifolia

Phenology Flowering and fruiting summer–fall. Flowering and fruiting summer–fall.
Habitat Disturbed areas. Stream banks, edges of woods, disturbed areas, agricultural fields.
Elevation 10–50 m. (0–200 ft.) 0–1700 m. (0–5600 ft.)
Distribution
from FNA
FL; Mexico; Central America; West Indies; n South America; c South America [Introduced in North America]
from FNA
AL; AR; AZ; FL; GA; IA; IL; IN; KS; KY; LA; MO; MS; NC; NE; OH; OK; SC; TN; TX; VA; WV; Mexico; West Indies [Introduced to Central America]
[WildflowerSearch map]
[BONAP county map]
Discussion

Acalypha arvensis became established in the United States in the 1980s and is now known from scattered localities throughout peninsular Florida. Some recent literature misapplies A. aristata Kunth to this species, but that name is a synonym of A. alopecuroidea.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

J. Torrey [in W. H. Emory 1857–1859, vol. 2(1)] reported Acalypha ostryifolia (as A. caroliniana Elliott) from New Mexico on the basis of Bigelow s.n., collected near "the Copper Mines" in Grant County. Although no specimen has been located by the author and this is the only report of this species from New Mexico, it is to be expected in the southwestern part of the state. It was collected in the late nineteenth century in New Jersey and Pennsylvania, but has not been collected there since.

Acalypha ostryifolia may not be native to the northern part of its range. Specimen collection dates suggest that the species is spreading northward, and in much of its range it is found primarily in areas with human disturbance.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 167. FNA vol. 12, p. 167.
Parent taxa Euphorbiaceae > Acalypha Euphorbiaceae > Acalypha
Sibling taxa
A. alopecuroidea, A. australis, A. californica, A. chamaedrifolia, A. deamii, A. gracilens, A. monococca, A. monostachya, A. neomexicana, A. ostryifolia, A. phleoides, A. poiretii, A. radians, A. rhomboidea, A. setosa, A. virginica, A. wilkesiana
A. alopecuroidea, A. arvensis, A. australis, A. californica, A. chamaedrifolia, A. deamii, A. gracilens, A. monococca, A. monostachya, A. neomexicana, A. phleoides, A. poiretii, A. radians, A. rhomboidea, A. setosa, A. virginica, A. wilkesiana
Name authority Poeppig: in E. F. Poeppig and S. L. Endlicher, Nov. Gen. Sp. Pl. 3: 21. (1841) Riddell ex J. M. Coulter: Mem. Torrey Bot. Club 5: 213. (1894) — (as ostryaefolia)
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