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Argentinian biddy-biddy, California sheepburr

Habit Plants from branched woody caudex.
Stems

decumbent, becoming suberect, 1.5–5 dm (in fruit), 4 mm diam.;

branches erect or leafy.

Leaves

(crowded at bases of stems);

stipules absent;

petiole to 4 cm;

blade 3–15 cm, leaflets 6–10 per side, gray-green (at least on dried specimens), ovate, 4–10 mm, pinnatisect, lobes 3–7(–11), linear, margins revolute, apex penicillate (having a little brush or tuft of hairs at tips), abaxial surface sericeous, adaxial sparsely pilose.

Scapes

terminal, to 50 cm, 1.5–3 mm diam., sparsely pilose, proximal scape and leaf sheaths with multicellular hairs among 1-celled hairs, with (0–)2–3 cauline leaves to 10 cm.

Inflorescences

15–35-flowered, interrupted spikes, (flowers crowded distally, 2–15 cm in flower, elongating to 50 cm in fruit);

bracteoles leaflike to linear, ciliate.

Flowers

6 mm diam.;

sepals 4–5, elliptic, 3 mm, abaxially pilose, adaxially glabrous;

stamens usually 4, filaments 2–3 mm, anthers dark red;

stigma rose, laciniate.

Fruiting

hypanthia ovoid, 3–4 mm diam., surfaces tomentose, obscurely 4-angled, bearing 10–20 spines in longitudinal ranks, to 3[–4] mm, spines with retrorse hairs or barbs.

Acaena pinnatifida

Phenology Flowering Mar–Jun.
Habitat Coastal grasslands, open rocky slopes, sand dunes, old roadsides
Elevation 50–400 m (200–1300 ft)
Distribution
from FNA
CA; South America (Argentina, Chile)
[WildflowerSearch map]
[BONAP county map]
Discussion

Herbarium records indicate occurrences of Acaena pinnatifida between Santa Barbara and Sonoma counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 324.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Agrimonieae > Acaena
Sibling taxa
A. novae-zelandiae, A. pallida
Synonyms A. californica, A. pinnatifida var. californica
Name authority Ruiz & Pavón: Fl. Peruv. 1: 68, plate 104, fig. b. (1798)
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