Acaena pinnatifida |
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Argentinian biddy-biddy, California sheepburr |
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Habit | Plants from branched woody caudex. |
Stems | decumbent, becoming suberect, 1.5–5 dm (in fruit), 4 mm diam.; branches erect or leafy. |
Leaves | (crowded at bases of stems); stipules absent; petiole to 4 cm; blade 3–15 cm, leaflets 6–10 per side, gray-green (at least on dried specimens), ovate, 4–10 mm, pinnatisect, lobes 3–7(–11), linear, margins revolute, apex penicillate (having a little brush or tuft of hairs at tips), abaxial surface sericeous, adaxial sparsely pilose. |
Scapes | terminal, to 50 cm, 1.5–3 mm diam., sparsely pilose, proximal scape and leaf sheaths with multicellular hairs among 1-celled hairs, with (0–)2–3 cauline leaves to 10 cm. |
Inflorescences | 15–35-flowered, interrupted spikes, (flowers crowded distally, 2–15 cm in flower, elongating to 50 cm in fruit); bracteoles leaflike to linear, ciliate. |
Flowers | 6 mm diam.; sepals 4–5, elliptic, 3 mm, abaxially pilose, adaxially glabrous; stamens usually 4, filaments 2–3 mm, anthers dark red; stigma rose, laciniate. |
Fruiting | hypanthia ovoid, 3–4 mm diam., surfaces tomentose, obscurely 4-angled, bearing 10–20 spines in longitudinal ranks, to 3[–4] mm, spines with retrorse hairs or barbs. |
Acaena pinnatifida |
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Phenology | Flowering Mar–Jun. |
Habitat | Coastal grasslands, open rocky slopes, sand dunes, old roadsides |
Elevation | 50–400 m (200–1300 ft) |
Distribution |
CA; South America (Argentina, Chile)
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Discussion | Herbarium records indicate occurrences of Acaena pinnatifida between Santa Barbara and Sonoma counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 324. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Agrimonieae > Acaena |
Sibling taxa | |
Synonyms | A. californica, A. pinnatifida var. californica |
Name authority | Ruiz & Pavón: Fl. Peruv. 1: 68, plate 104, fig. b. (1798) |
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