Abronia fragrans |
Abronia umbellata |
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fragrant verbena, fragrant white sand-verbena, heart's-delight, snowball sand-verbena |
beach sand verbena, pink sand-verbena, purple sand verbena |
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Habit | Plants perennial. | Plants annual. | ||||||||
Stems | procumbent to semierect, slightly to moderately branched, elongate, sometimes reddish at base and nodes, glandular-pubescent, viscid. |
prostrate, much branched in large plants, forming loose mats, elongate, glandular-pubescent or glabrous. |
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Leaves | petiole 0.5–8 cm; blade ovate to triangular or lanceolate, 3–12 × 1–8 cm, margins entire to subsinuate and slightly undulate, adaxial surface glandular-pubescent, abaxial surface more densely and longer pubescent, or sometimes villous. |
petiole 1–6 cm; blade ovate, elliptic, or rhombic, 1.5–6.8 × 0.8–4.7 cm, margins entire to ± repand and undulate, surfaces glandular-puberulent to glandular-villous, usually ± glaucous. |
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Inflorescences | peduncle longer than subtending petiole; bracts linear-lanceolate to oval-ovate, 7–25 × 2–12 mm, scarious, glandular-puberulent to short villous; flowers 30–80. |
peduncle longer than subtending petiole; bracts lanceolate to ovate, 5–7 × 3–5 mm, papery, glandular-puberulent to viscid-villous; flowers 8–27. |
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Perianth | tube greenish to reddish purple, 10–25 mm, limb white, (2–)6–10 mm diam. |
tube magenta to greenish, 6.5–18 mm, limb magenta, 6–16 mm diam., throat surrounded by white to yellowish white eyespot. |
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Fruits | winged or not, fusiform and appearing deeply grooved when wingless, when wings not distorted, fruit ± cordate in profile, tapered at base, with prominent beak in broad notch at apex, 5–12 × 2.5–7 mm, indurate, rugose veined or, if wings distorted, fruit ± rhombic in profile and tapered at both ends, or obdeltate and truncate at apex, with prominent beak; wings 4–5, thick, narrow, not dilated at apex, cavities extending throughout; peripheral fruits often distorted, S-shaped in lateral view. |
± obdeltate in profile, 6–12 × 6–16(–24) mm, indurate, smooth, not rugose veined, apex beaklike; wings 5, not folded, poorly to very well developed, from slightly shorter than beaklike apex of body to prolonged beyond beak, thin, without cavities. |
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Abronia fragrans |
Abronia umbellata |
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Phenology | Flowering spring–fall. | |||||||||
Habitat | Dry sandy soils, scrub and grasslands | |||||||||
Elevation | 400-2000 m (1300-6600 ft) | |||||||||
Distribution |
AZ; CO; KS; MT; ND; NE; NM; OK; SD; TX; UT; WY; Mexico (Chihuahua)
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CA; OR; WA; BC; Mexico (Baja California)
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Discussion | Varieties 3 (3 in the flora). S. S. Tillett (1967) stated that Abronia umbellata is sufficiently similar to the inland A. villosa that without information regarding locality some specimens would be very difficult to identify. He also considered A. umbellata subsp. alba (Eastwood) Munz, subsp. platyphylla (Standley) Munz, and subsp. variabilis (Standley) Munz, A. insularis Standley, and A. neurophylla Standley to be introgressive hybrids of A. umbellata with A. maritima. The name Abronia gracilis Bentham has appeared in regional floras and treatments of Nyctaginaceae since Standley’s continental treatment (1918). S. S. Tillett (1967) stated that the taxon was reported for San Diego County, California, but cited no documenting specimens. I. L. Wiggins (F. Shreve and I. L. Wiggins 1964) considered only A. gracilis subsp. platyphylla (Standley) Ferris to enter the United States, in San Diego County, an entity considered by Tillett to be an intergrade between A. umbellata and A. maritima. The distinctions among A. gracilis, A. umbellata, and A. villosa are subtle at best, and the group is in need of careful study. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 4, p. 63. | FNA vol. 4, p. 67. | ||||||||
Parent taxa | Nyctaginaceae > Abronia | Nyctaginaceae > Abronia | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Name authority | Nuttall ex Hooker: Hooker’s J. Bot. Kew Gard. Misc. 5: 261. (1853) | Lamarck: Tabl. Encycl. 1: 469, plate 105. (1791) | ||||||||
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