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fragrant verbena, fragrant white sand-verbena, heart's-delight, snowball sand-verbena

dwarf sand verbena

Habit Plants perennial. Plants perennial, acaulescent or nearly so, usually cespitose.
Stems

procumbent to semierect, slightly to moderately branched, elongate, sometimes reddish at base and nodes, glandular-pubescent, viscid.

Leaves

petiole 0.5–8 cm;

blade ovate to triangular or lanceolate, 3–12 × 1–8 cm, margins entire to subsinuate and slightly undulate, adaxial surface glandular-pubescent, abaxial surface more densely and longer pubescent, or sometimes villous.

petiole 1–5 cm;

blade elliptic to lanceolate, shortly ovate, or oblong-ovate, (0.4–)0.5–2.5 × (0.2–)0.4–1.2 cm, less than 3 times as long as wide, margins entire or ± repand and undulate, surfaces glabrous or glandular-pubescent.

Inflorescences

peduncle longer than subtending petiole;

bracts linear-lanceolate to oval-ovate, 7–25 × 2–12 mm, scarious, glandular-puberulent to short villous;

flowers 30–80.

bracts lanceolate to ovate, 4–9 × 2–7 mm, scarious, glandular-puberulent, often minutely so;

flowers 15–25.

Perianth

tube greenish to reddish purple, 10–25 mm, limb white, (2–)6–10 mm diam.

tube pale pink, 8–30 mm, limb white to pink, 6–10 mm diam.

Fruits

winged or not, fusiform and appearing deeply grooved when wingless, when wings not distorted, fruit ± cordate in profile, tapered at base, with prominent beak in broad notch at apex, 5–12 × 2.5–7 mm, indurate, rugose veined or, if wings distorted, fruit ± rhombic in profile and tapered at both ends, or obdeltate and truncate at apex, with prominent beak;

wings 4–5, thick, narrow, not dilated at apex, cavities extending throughout;

peripheral fruits often distorted, S-shaped in lateral view.

obovate to obcordate in profile, 6–10 × 5–7 mm, scarious, apex low and broadly conic;

wings 5, without dilations, without cavities.

Abronia fragrans

Abronia nana

Phenology Flowering spring–fall.
Habitat Dry sandy soils, scrub and grasslands
Elevation 400-2000 m (1300-6600 ft)
Distribution
from FNA
AZ; CO; KS; MT; ND; NE; NM; OK; SD; TX; UT; WY; Mexico (Chihuahua)
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; NV; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Abronia nana is a highly variable species. Perhaps contraction of the range of A. nana during the Pleistocene left isolated populations that have since diverged. This is especially apparent on the southern edge of the range of the species. In northeastern Arizona, densely tufted plants with very small blades are similar to short-leaved plants of A. bigelovii from north-central New Mexico.

Based on the fruits, the taxon described as Abronia nana var. harrisii S. L. Welsh is A. elliptica.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades elliptic-lanceolate to elliptic-ovate; inflorescence bracts ovate to oblong-lanceolate, or vestigial
var. nana
1. Leaf blades shortly ovate to oblong-ovate; inflorescence bracts lanceolate
var. covillei
Source FNA vol. 4, p. 63. FNA vol. 4, p. 69.
Parent taxa Nyctaginaceae > Abronia Nyctaginaceae > Abronia
Sibling taxa
A. alpina, A. ameliae, A. ammophila, A. angustifolia, A. argillosa, A. bigelovii, A. bolackii, A. carletonii, A. elliptica, A. latifolia, A. macrocarpa, A. maritima, A. mellifera, A. nana, A. pogonantha, A. turbinata, A. umbellata, A. villosa
A. alpina, A. ameliae, A. ammophila, A. angustifolia, A. argillosa, A. bigelovii, A. bolackii, A. carletonii, A. elliptica, A. fragrans, A. latifolia, A. macrocarpa, A. maritima, A. mellifera, A. pogonantha, A. turbinata, A. umbellata, A. villosa
Subordinate taxa
A. nana var. covillei, A. nana var. nana
Name authority Nuttall ex Hooker: Hooker’s J. Bot. Kew Gard. Misc. 5: 261. (1853) S. Watson: Proc. Amer. Acad. Arts. 14: 294. (1879)
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