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fragrant verbena, fragrant white sand-verbena, heart's-delight, snowball sand-verbena

Wyoming sand verbena

Habit Plants perennial. Plants perennial.
Stems

procumbent to semierect, slightly to moderately branched, elongate, sometimes reddish at base and nodes, glandular-pubescent, viscid.

prostrate, much branched, forming loose mats, elongate, densely glandular-pubescent, viscid.

Leaves

petiole 0.5–8 cm;

blade ovate to triangular or lanceolate, 3–12 × 1–8 cm, margins entire to subsinuate and slightly undulate, adaxial surface glandular-pubescent, abaxial surface more densely and longer pubescent, or sometimes villous.

petiole 2–4.5 cm;

blade elliptic-oblong to rhombic-ovate, 1–2.5 × 0.6–1.5 cm, margins entire to repand and ± undulate, surfaces glandular-puberulent.

Inflorescences

peduncle longer than subtending petiole;

bracts linear-lanceolate to oval-ovate, 7–25 × 2–12 mm, scarious, glandular-puberulent to short villous;

flowers 30–80.

peduncle longer than subtending petiole;

bracts lanceolate to elliptic, 3–5 × 1–3 mm, papery, puberulent to glandular-pubescent;

flowers 15–35.

Perianth

tube greenish to reddish purple, 10–25 mm, limb white, (2–)6–10 mm diam.

tube greenish, 7–12 mm, limb greenish white, 4–6 mm diam.

Fruits

winged or not, fusiform and appearing deeply grooved when wingless, when wings not distorted, fruit ± cordate in profile, tapered at base, with prominent beak in broad notch at apex, 5–12 × 2.5–7 mm, indurate, rugose veined or, if wings distorted, fruit ± rhombic in profile and tapered at both ends, or obdeltate and truncate at apex, with prominent beak;

wings 4–5, thick, narrow, not dilated at apex, cavities extending throughout;

peripheral fruits often distorted, S-shaped in lateral view.

biturbinate, tapered at both ends, ± rhombic or fusiform in profile, 4–6 × 2.5–4.5 mm, indurate;

wings reduced, appearing as 5 prominent lobes near middle of fruit;

peripheral fruits not or only slightly distorted.

Abronia fragrans

Abronia ammophila

Phenology Flowering spring–fall. Flowering summer–fall.
Habitat Dry sandy soils, scrub and grasslands Sandy soils, lake shores
Elevation 400-2000 m (1300-6600 ft) 2300-2500 m (7500-8200 ft)
Distribution
from FNA
AZ; CO; KS; MT; ND; NE; NM; OK; SD; TX; UT; WY; Mexico (Chihuahua)
[WildflowerSearch map]
[BONAP county map]
from FNA
WY
[BONAP county map]
Discussion

Of conservation concern.

Abronia ammophila is found on the beaches of Yellowstone Lake, Yellowstone National Park.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 4, p. 63. FNA vol. 4, p. 67.
Parent taxa Nyctaginaceae > Abronia Nyctaginaceae > Abronia
Sibling taxa
A. alpina, A. ameliae, A. ammophila, A. angustifolia, A. argillosa, A. bigelovii, A. bolackii, A. carletonii, A. elliptica, A. latifolia, A. macrocarpa, A. maritima, A. mellifera, A. nana, A. pogonantha, A. turbinata, A. umbellata, A. villosa
A. alpina, A. ameliae, A. angustifolia, A. argillosa, A. bigelovii, A. bolackii, A. carletonii, A. elliptica, A. fragrans, A. latifolia, A. macrocarpa, A. maritima, A. mellifera, A. nana, A. pogonantha, A. turbinata, A. umbellata, A. villosa
Synonyms A. arenaria
Name authority Nuttall ex Hooker: Hooker’s J. Bot. Kew Gard. Misc. 5: 261. (1853) Greene: Pittonia 4: 226. (1900)
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