Burke: Penstemon pruinosus vs. Penstemon washingtonensis

	Penstemon pruinosus	Penstemon washingtonensis
	Chelan beardtongue	Washington beardtongue
Habit	Plants tufted from a stout, branched, and woody base, the stems 1-4 dm. tall; inflorescence always glandular-hairy, the rest of the plant varying from glandular-hairy to glabrous.	Tufted perennials from a loose, woody rhizome, the stems 1-2.5 dm. tall, usually glabrous below the glandular-hairy inflorescence.
Leaves	Leaves opposite, usually sharply serrate, sometimes most of them entire; basal leaves well-developed, up to 10 cm. long and 2 cm. wide, the blades elliptic to ovate, equaling the petiole; cauline leaves sessile, triangular-ovate to narrowly lanceolate, smaller than the basal leaves.	Leaves opposite, entire, the basal ones well-developed and forming rosettes, 2.5-6 cm. long and 5-18 mm. wide; cauline leaves few, sessile, often reduced.
Flowers	Inflorescence of several compact verticillasters; calyx 3-6 mm. long, the 5 segments broadly lanceolate; corolla deep blue or lavender, 11-16 mm. long, the tube slightly expanded, 2.5-5 mm. wide at the mouth, the throat paler and marked with guide lines, glandular-hairy externally, the raised portion of the lower lip bearded; pollen sacs glabrous, dehiscent throughout and becoming opposite; staminode with a bearded tip.	Inflorescence of 1-3 dense verticillasters; calyx 4-6 mm. long, the 5 segments tapered, with scarious margins; corolla deep blue or occasionally pale yellow, glandular-hairy, 9-12 mm. long, the tube narrow, only 2-3 mm. wide at the mouth, the raised portion of the lower lip bearded; pollen sacs glabrous, sub-rotund, 0.5-0.6 mm. long, wholly dehiscent, becoming opposite; staminode bearded toward the expanded tip.
Fruits	Capsule 5-7 mm. long, glabrous.	Capsule 4-5 mm. long

	Penstemon pruinosus	Penstemon washingtonensis
Flowering time	May-July	June-August
Habitat	Open, rocky places from the valleys and plains to moderate elevations in the mountains, and in the scablands in Washington.	From open slopes and flats at moderate elevations to alpine meadows.
Distribution	Occurring east of the Cascades crest in Washington; British Columbia to Washington. [WildflowerSearch map] [BONAP county map]	Occurring east of the Cascades crest, where endemic to Chelan and Okanogan in Washington. [WildflowerSearch map] [BONAP county map]
Origin	Native	Native
Conservation status	Not of concern	Not of concern
Sibling taxa	P. acuminatus, P. attenuatus, P. barrettiae, P. cardwellii, P. cinereus, P. confertus, P. davidsonii, P. deustus, P. ellipticus, P. eriantherus, P. euglaucus, P. fruticosus, P. gairdneri, P. glandulosus, P. hesperius, P. lyallii, P. ovatus, P. palmeri, P. pennellianus, P. procerus, P. richardsonii, P. rupicola, P. rydbergii, P. serrulatus, P. speciosus, P. subserratus, P. triphyllus, P. venustus, P. washingtonensis, P. wilcoxii	P. acuminatus, P. attenuatus, P. barrettiae, P. cardwellii, P. cinereus, P. confertus, P. davidsonii, P. deustus, P. ellipticus, P. eriantherus, P. euglaucus, P. fruticosus, P. gairdneri, P. glandulosus, P. hesperius, P. lyallii, P. ovatus, P. palmeri, P. pennellianus, P. procerus, P. pruinosus, P. richardsonii, P. rupicola, P. rydbergii, P. serrulatus, P. speciosus, P. subserratus, P. triphyllus, P. venustus, P. wilcoxii
Web links	Local floras: BC, WA Local Web sites: Flora NW, PNW Herbaria, Turner Photog. WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: WA Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Penstemon washingtonensis

Chelan beardtongue

Washington beardtongue

Habit

Plants tufted from a stout, branched, and woody base, the stems 1-4 dm. tall; inflorescence always glandular-hairy, the rest of the plant varying from glandular-hairy to glabrous.

Tufted perennials from a loose, woody rhizome, the stems 1-2.5 dm. tall, usually glabrous below the glandular-hairy inflorescence.

Leaves

Leaves opposite, usually sharply serrate, sometimes most of them entire;

basal leaves well-developed, up to 10 cm. long and 2 cm. wide, the blades elliptic to ovate, equaling the petiole;

cauline leaves sessile, triangular-ovate to narrowly lanceolate, smaller than the basal leaves.

Leaves opposite, entire, the basal ones well-developed and forming rosettes, 2.5-6 cm. long and 5-18 mm. wide;

cauline leaves few, sessile, often reduced.

Flowers

Inflorescence of several compact verticillasters;

calyx 3-6 mm. long, the 5 segments broadly lanceolate;

corolla deep blue or lavender, 11-16 mm. long, the tube slightly expanded, 2.5-5 mm. wide at the mouth, the throat paler and marked with guide lines, glandular-hairy externally, the raised portion of the lower lip bearded;

pollen sacs glabrous, dehiscent throughout and becoming opposite;

staminode with a bearded tip.

Inflorescence of 1-3 dense verticillasters;

calyx 4-6 mm. long, the 5 segments tapered, with scarious margins;

corolla deep blue or occasionally pale yellow, glandular-hairy, 9-12 mm. long, the tube narrow, only 2-3 mm. wide at the mouth, the raised portion of the lower lip bearded;

pollen sacs glabrous, sub-rotund, 0.5-0.6 mm. long, wholly dehiscent, becoming opposite;

staminode bearded toward the expanded tip.

Fruits

Capsule 5-7 mm. long, glabrous.

Capsule 4-5 mm. long