Burke: Penstemon harbourii vs. Penstemon wilcoxii

	Penstemon harbourii	Penstemon wilcoxii
		Wilcox's beardtongue
Habit		Perennial from a branched, woody base, the clustered stems 3-10 dm. tall, usually glabrous except for the inflorescence
Leaves		Leaves opposite, strongly serrate with slender teeth, the basal leaves well-developed, elliptic to deltoid-ovate, the blade up to 9 cm. long and 5 cm. wide, about equal to the petiole; cauline leaves often larger than the basal, sessile, 4-10 cm. long and 1.5-4 cm. wide.
Flowers		Inflorescence of several large, loose verticillasters; calyx 2.5-5 mm. long, the 5 segments broadly lanceolate, narrowly scarious-margined; corolla blue, the paler throat with guide lines, 15-23 mm. long, the tube 4-8 mm. wide at the mouth, glandular-hairy on the outside, strongly bilabiate, the lower lip much longer than the upper, the raised portion bearded; pollen sacs 0.7-1.0 mm. long, wholly dehiscent, becoming opposite, glabrous; staminode with a recurved, bearded tip.
Fruits		Capsule 4-6 mm. long.

	Penstemon harbourii	Penstemon wilcoxii
Flowering time		May-July
Habitat		Open or often wooded, sometimes in rocky places, from the foothills to moderate elevations in the mountains.
Distribution	[WildflowerSearch map] [BONAP county map]	Occurring east of the Cascades crest in eastern Washington; Washington to Oregon, east to Montana. [WildflowerSearch map] [BONAP county map]
Origin		Native
Conservation status		Threatened in Washington (WANHP)
Sibling taxa	P. acuminatus, P. attenuatus, P. barrettiae, P. cardwellii, P. cinereus, P. confertus, P. davidsonii, P. deustus, P. ellipticus, P. eriantherus, P. euglaucus, P. fruticosus, P. gairdneri, P. glandulosus, P. hesperius, P. lyallii, P. ovatus, P. palmeri, P. pennellianus, P. procerus, P. pruinosus, P. richardsonii, P. rupicola, P. rydbergii, P. serrulatus, P. speciosus, P. subserratus, P. triphyllus, P. venustus, P. washingtonensis, P. wilcoxii	P. acuminatus, P. attenuatus, P. barrettiae, P. cardwellii, P. cinereus, P. confertus, P. davidsonii, P. deustus, P. ellipticus, P. eriantherus, P. euglaucus, P. fruticosus, P. gairdneri, P. glandulosus, P. hesperius, P. lyallii, P. ovatus, P. palmeri, P. pennellianus, P. procerus, P. pruinosus, P. richardsonii, P. rupicola, P. rydbergii, P. serrulatus, P. speciosus, P. subserratus, P. triphyllus, P. venustus, P. washingtonensis
Web links	Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: OR, WA Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Penstemon harbourii

Penstemon wilcoxii

Wilcox's beardtongue

Habit

Perennial from a branched, woody base, the clustered stems 3-10 dm. tall, usually glabrous except for the inflorescence

Leaves

Leaves opposite, strongly serrate with slender teeth, the basal leaves well-developed, elliptic to deltoid-ovate, the blade up to 9 cm. long and 5 cm. wide, about equal to the petiole;

cauline leaves often larger than the basal, sessile, 4-10 cm. long and 1.5-4 cm. wide.

Flowers

Inflorescence of several large, loose verticillasters;

calyx 2.5-5 mm. long, the 5 segments broadly lanceolate, narrowly scarious-margined;

corolla blue, the paler throat with guide lines, 15-23 mm. long, the tube 4-8 mm. wide at the mouth, glandular-hairy on the outside, strongly bilabiate, the lower lip much longer than the upper, the raised portion bearded;

pollen sacs 0.7-1.0 mm. long, wholly dehiscent, becoming opposite, glabrous;

staminode with a recurved, bearded tip.

Fruits

Capsule 4-6 mm. long.