Burke: Penstemon harbourii vs. Penstemon washingtonensis

	Penstemon harbourii	Penstemon washingtonensis
		Washington beardtongue
Habit		Tufted perennials from a loose, woody rhizome, the stems 1-2.5 dm. tall, usually glabrous below the glandular-hairy inflorescence.
Leaves		Leaves opposite, entire, the basal ones well-developed and forming rosettes, 2.5-6 cm. long and 5-18 mm. wide; cauline leaves few, sessile, often reduced.
Flowers		Inflorescence of 1-3 dense verticillasters; calyx 4-6 mm. long, the 5 segments tapered, with scarious margins; corolla deep blue or occasionally pale yellow, glandular-hairy, 9-12 mm. long, the tube narrow, only 2-3 mm. wide at the mouth, the raised portion of the lower lip bearded; pollen sacs glabrous, sub-rotund, 0.5-0.6 mm. long, wholly dehiscent, becoming opposite; staminode bearded toward the expanded tip.
Fruits		Capsule 4-5 mm. long

	Penstemon harbourii	Penstemon washingtonensis
Flowering time		June-August
Habitat		From open slopes and flats at moderate elevations to alpine meadows.
Distribution	[WildflowerSearch map] [BONAP county map]	Occurring east of the Cascades crest, where endemic to Chelan and Okanogan in Washington. [WildflowerSearch map] [BONAP county map]
Origin		Native
Conservation status		Not of concern
Sibling taxa	P. acuminatus, P. attenuatus, P. barrettiae, P. cardwellii, P. cinereus, P. confertus, P. davidsonii, P. deustus, P. ellipticus, P. eriantherus, P. euglaucus, P. fruticosus, P. gairdneri, P. glandulosus, P. hesperius, P. lyallii, P. ovatus, P. palmeri, P. pennellianus, P. procerus, P. pruinosus, P. richardsonii, P. rupicola, P. rydbergii, P. serrulatus, P. speciosus, P. subserratus, P. triphyllus, P. venustus, P. washingtonensis, P. wilcoxii	P. acuminatus, P. attenuatus, P. barrettiae, P. cardwellii, P. cinereus, P. confertus, P. davidsonii, P. deustus, P. ellipticus, P. eriantherus, P. euglaucus, P. fruticosus, P. gairdneri, P. glandulosus, P. hesperius, P. lyallii, P. ovatus, P. palmeri, P. pennellianus, P. procerus, P. pruinosus, P. richardsonii, P. rupicola, P. rydbergii, P. serrulatus, P. speciosus, P. subserratus, P. triphyllus, P. venustus, P. wilcoxii
Web links	Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: WA Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Penstemon harbourii

Penstemon washingtonensis

Washington beardtongue

Habit

Tufted perennials from a loose, woody rhizome, the stems 1-2.5 dm. tall, usually glabrous below the glandular-hairy inflorescence.

Leaves

Leaves opposite, entire, the basal ones well-developed and forming rosettes, 2.5-6 cm. long and 5-18 mm. wide;

cauline leaves few, sessile, often reduced.

Flowers

Inflorescence of 1-3 dense verticillasters;

calyx 4-6 mm. long, the 5 segments tapered, with scarious margins;

corolla deep blue or occasionally pale yellow, glandular-hairy, 9-12 mm. long, the tube narrow, only 2-3 mm. wide at the mouth, the raised portion of the lower lip bearded;

pollen sacs glabrous, sub-rotund, 0.5-0.6 mm. long, wholly dehiscent, becoming opposite;

staminode bearded toward the expanded tip.

Fruits

Capsule 4-5 mm. long