Burke: Penstemon glandulosus vs. Penstemon barrettiae

	Penstemon glandulosus	Penstemon barrettiae
	glandular penstemon	Barrett's beardtongue
Habit	Stout perennial herb from a branched, woody base, the several stems 4-10 dm. tall, sticky throughout with stalked glands.	Perennial, the lower part shrubby and branched, the stems 2-4 dm. tall; herbage and inflorescence glabrous and glaucous.
Leaves	Leaves opposite, sharply toothed or entire, the basal ones 10-35 cm. long and 2.5-9 cm. wide, petiolate, the blade lanceolate to elliptic; lowermost cauline leaves reduced, the others well-developed, sessile and often clasping, broadly lanceolate to rotund-ovate, 4-12 cm. long and 2-5 cm. wide.	Leaves opposite, firm, irregularly serrulate or entire, the larger ones, up to 8 cm. long and 2.5 cm. wide, on sterile shoots borne toward the base of the main stem; cauline leaves broad, sessile, and clasping, 1.5-3.5 cm. long and 0.8-2 cm. wide
Flowers	Inflorescence of several verticillasters; calyx 9-15 mm. long, the 5 segments narrow and herbaceous; corolla blue-lavender, strongly glandular-hairy outside, glabrous inside, 28-40 mm. long, over 1 cm. wide at the mouth; anthers permanently horseshoe-shaped, 1.7-2.3 mm. long; pollen sacs dehiscent across their apices, the lower part saccate and indehiscent, glabrous; staminode glabrous, the tip flattened.	Inflorescence essentially a raceme, the axillary peduncles simple and single-flowered or sometimes branched and 2-flowered; calyx 5-7 mm. long, the 5 segments thin, ovate; corolla bilabiate, lilac or rose-purple, 33-38 mm. long, 1 cm. wide at the mouth, glabrous outside, long-hairy near the base of the lower lip within; anthers densely long-woolly, pollen sacs wholly dehiscent, becoming opposite
Fruits	Capsules 10-14 mm. long, surpassed by the calyx.	Capsule narrow, up to 1 cm. long.

	Penstemon glandulosus	Penstemon barrettiae
Flowering time	May-July	April-May
Habitat	Open, often rocky hillsides and banks in the foothills, valleys, and lower mountains.	Woodland openings and rocky slopes at low elevations.
Distribution	Occurring east of the Cascades crest in Washington; central Washington to north-central Oregon, and from southeastern Washington to eastern Oregon, east to Idaho. [WildflowerSearch map] [BONAP county map]	Occurring in the east end of the Columbia River Gorge in Washington; south-central Washington to adjacent north-central Oregon. [WildflowerSearch map] [BONAP county map]
Origin	Native	Native
Conservation status	Not of concern	Threatened in Washington (WANHP)
Sibling taxa	P. acuminatus, P. attenuatus, P. barrettiae, P. cardwellii, P. cinereus, P. confertus, P. davidsonii, P. deustus, P. ellipticus, P. eriantherus, P. euglaucus, P. fruticosus, P. gairdneri, P. hesperius, P. lyallii, P. ovatus, P. palmeri, P. pennellianus, P. procerus, P. pruinosus, P. richardsonii, P. rupicola, P. rydbergii, P. serrulatus, P. speciosus, P. subserratus, P. triphyllus, P. venustus, P. washingtonensis, P. wilcoxii	P. acuminatus, P. attenuatus, P. cardwellii, P. cinereus, P. confertus, P. davidsonii, P. deustus, P. ellipticus, P. eriantherus, P. euglaucus, P. fruticosus, P. gairdneri, P. glandulosus, P. hesperius, P. lyallii, P. ovatus, P. palmeri, P. pennellianus, P. procerus, P. pruinosus, P. richardsonii, P. rupicola, P. rydbergii, P. serrulatus, P. speciosus, P. subserratus, P. triphyllus, P. venustus, P. washingtonensis, P. wilcoxii
Subordinate taxa	P. glandulosus var. chelanensis, P. glandulosus var. glandulosus
Web links	Local floras: OR, WA Local Web sites: Flora NW, PNW Herbaria, Turner Photog. WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: OR, WA Local Web sites: Flora NW, PNW Herbaria, Turner Photog. WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Penstemon barrettiae

glandular penstemon

Barrett's beardtongue

Habit

Stout perennial herb from a branched, woody base, the several stems 4-10 dm. tall, sticky throughout with stalked glands.

Perennial, the lower part shrubby and branched, the stems 2-4 dm. tall; herbage and inflorescence glabrous and glaucous.

Leaves

Leaves opposite, sharply toothed or entire, the basal ones 10-35 cm. long and 2.5-9 cm. wide, petiolate, the blade lanceolate to elliptic; lowermost cauline leaves reduced, the others well-developed, sessile and often clasping, broadly lanceolate to rotund-ovate, 4-12 cm. long and 2-5 cm. wide.

Leaves opposite, firm, irregularly serrulate or entire, the larger ones, up to 8 cm. long and 2.5 cm. wide, on sterile shoots borne toward the base of the main stem;

cauline leaves broad, sessile, and clasping, 1.5-3.5 cm. long and 0.8-2 cm. wide

Flowers

Inflorescence of several verticillasters;

calyx 9-15 mm. long, the 5 segments narrow and herbaceous;

corolla blue-lavender, strongly glandular-hairy outside, glabrous inside, 28-40 mm. long, over 1 cm. wide at the mouth;

anthers permanently horseshoe-shaped, 1.7-2.3 mm. long;

pollen sacs dehiscent across their apices, the lower part saccate and indehiscent, glabrous;

staminode glabrous, the tip flattened.

Inflorescence essentially a raceme, the axillary peduncles simple and single-flowered or sometimes branched and 2-flowered;

calyx 5-7 mm. long, the 5 segments thin, ovate;

corolla bilabiate, lilac or rose-purple, 33-38 mm. long, 1 cm. wide at the mouth, glabrous outside, long-hairy near the base of the lower lip within;

anthers densely long-woolly, pollen sacs wholly dehiscent, becoming opposite

Fruits

Capsules 10-14 mm. long, surpassed by the calyx.

Capsule narrow, up to 1 cm. long.

Penstemon glandulosus

Penstemon barrettiae

Flowering time

May-July

April-May

Habitat

Open, often rocky hillsides and banks in the foothills, valleys, and lower mountains.

Woodland openings and rocky slopes at low elevations.

Distribution

Occurring east of the Cascades crest in Washington; central Washington to north-central Oregon, and from southeastern Washington to eastern Oregon, east to Idaho.

[WildflowerSearch map]

[BONAP county map]