Burke: Penstemon dolius vs. Penstemon cardwellii

	Penstemon dolius	Penstemon cardwellii
		Cardwell's beardtongue
Habit		Low shrub, the stems 1-3 dm. tall, ascending or erect.
Leaves		Leaves opposite, glabrous, the largest on short, sterile shoots, serrulate or nearly entire, short-petiolate, the blades elliptic, 1.5-3.5 cm. long and 6-14 mm. wide; leaves of the flowering shoots less crowded, smaller, sessile and entire.
Flowers		Inflorescence racemose, few-flowered and crowded, glandular, the simple pedicels opposite and axillary; calyx 5-12 mm. long, the 5 segments thin and lanceolate; corolla bright purple to deep blue-violet, 30-38 mm. long, about 1 cm. wide at the mouth, keeled on the back, glabrous outside and with long, white hairs near the base of the lower lip within; anthers long-wooly, pollen sacs opposite; staminode slender, shorter than the 4 fertile filaments, long-bearded toward the tip
Fruits		Capsule

	Penstemon dolius	Penstemon cardwellii
Flowering time		May-July
Habitat		Open or wooded summits or slopes at middle elevations in the mountains.
Distribution	[WildflowerSearch map] [BONAP county map]	Occurring west of the Cascades crest in Washington; southwestern Washington to southwestern Oregon. [WildflowerSearch map] [BONAP county map]
Origin		Native
Conservation status		Not of concern
Sibling taxa	P. acuminatus, P. attenuatus, P. barrettiae, P. cardwellii, P. cinereus, P. confertus, P. davidsonii, P. deustus, P. ellipticus, P. eriantherus, P. euglaucus, P. fruticosus, P. gairdneri, P. glandulosus, P. hesperius, P. lyallii, P. ovatus, P. palmeri, P. pennellianus, P. procerus, P. pruinosus, P. richardsonii, P. rupicola, P. rydbergii, P. serrulatus, P. speciosus, P. subserratus, P. triphyllus, P. venustus, P. washingtonensis, P. wilcoxii	P. acuminatus, P. attenuatus, P. barrettiae, P. cinereus, P. confertus, P. davidsonii, P. deustus, P. ellipticus, P. eriantherus, P. euglaucus, P. fruticosus, P. gairdneri, P. glandulosus, P. hesperius, P. lyallii, P. ovatus, P. palmeri, P. pennellianus, P. procerus, P. pruinosus, P. richardsonii, P. rupicola, P. rydbergii, P. serrulatus, P. speciosus, P. subserratus, P. triphyllus, P. venustus, P. washingtonensis, P. wilcoxii
Web links	Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: OR, WA Local Web sites: Flora NW, PNW Herbaria, Turner Photog. WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Penstemon dolius

Penstemon cardwellii

Cardwell's beardtongue

Habit

Low shrub, the stems 1-3 dm. tall, ascending or erect.

Leaves

Leaves opposite, glabrous, the largest on short, sterile shoots, serrulate or nearly entire, short-petiolate, the blades elliptic, 1.5-3.5 cm. long and 6-14 mm. wide;

leaves of the flowering shoots less crowded, smaller, sessile and entire.

Flowers

Inflorescence racemose, few-flowered and crowded, glandular, the simple pedicels opposite and axillary;

calyx 5-12 mm. long, the 5 segments thin and lanceolate;

corolla bright purple to deep blue-violet, 30-38 mm. long, about 1 cm. wide at the mouth, keeled on the back, glabrous outside and with long, white hairs near the base of the lower lip within;

anthers long-wooly, pollen sacs opposite;

staminode slender, shorter than the 4 fertile filaments, long-bearded toward the tip

Fruits

Capsule