Burke: Erythranthe washingtonensis vs. Erythranthe cardinalis

	Erythranthe washingtonensis	Erythranthe cardinalis
	Washington monkey-flower	scarlet monkey-flower
Habit	Annuals with fibrous roots or threadlike taproot; stems 5-25 cm, erect to ascending, straight or sharply bent at nodes, typically heavily branched, not angled; puberulent- to villous-glandular, flattened gland-tipped hairs 0.1-0.8 mm, sometimes transparent.	Rhizomatous perennials; stems 25-75 cm, usually erect or curving upward, branched, somewhat glandular-villous or becoming glabrous.
Leaves	Leaves cauline, basal ones not persistent; petioles 2-14 mm; blade triangular to ovate to narrowly ovate, 4-16 mm long and 2-11 mm broad, palmate venation, base rounded to cuneate to truncate, margins finely toothed or entire, apex acute, surfaces hairy-glandular as stems.	Leaves typically cauline; sessile; blade ovate to elliptic to obovate, 20-90 mm long and 10-38 mm broad, palmate venation, base acuminate and nearly clasping, margins irregularly toothed, apex acute, surfaces hairy as stems.
Flowers	Axillary flowers 1-6, emerging from nodes throughout; fruiting pedicels 20-50 mm, densely covered with minute stalked glands; calyx greenish, ridged, tubular, slightly inflated, 6-8 mm, margins toothed or lobed, covered with minute stalked glands, lobes pronounced, erect; corollas yellow with small reddish brown dots, lower limb with two whitish patches, symmetric bilaterally, bilabiate; tube-throat funnel-shaped and 8-10 mm, protruding beyond calyx margin; limb expanded 7-10 mm, lobes obovate-oblong, apex rounded to somewhat cuneate; styles hispid-hirtellous; anthers not protruding, glabrous.	Axillary flowers 2-12, emerging from leaf axes at mid- to ends of stem; fruiting pedicels generally 30-90 mm; calyx cylindric to nearly bell-shaped, not inflated, 17-28 mm, hairy or bristly-hairy, lobes 4-7 mm, ovate to nearly deltate, apex attenuate-acute; corollas red to orangish red or rarely yellow, throat yellowish with red stripes, palate red, yellow-villous and not spotted or striped, symmetric bilaterally, distinctly bilabiate; tube-throat funnel-shaped, generally 20-30 mm, protruding beyond calyx margin; throat open; styles glabrous; anthers protruding, white-hairy, thecae spreading.
Fruits	Capsules 5-8.5 mm, included.	Capsules 10-16 mm, included.

	Erythranthe washingtonensis	Erythranthe cardinalis
Flowering time	April-June	May-August
Habitat	Shallow or gravelly soils of vernally moist areas at low elevations.	Streambanks, where escaped from cultivation.
Distribution	Occurring east of the cascades crest in the Columbia River Gorge in Washington; south-central Washington to adjacent north-central Oregon. [WildflowerSearch map]	Occurring east of the Cascades crest in Washington; Washington to California, east to the southern Rocky Mountains. [WildflowerSearch map]
Origin	Native	Introduced
Conservation status	Historical in Washington (WANHP)	Not of concern
Sibling taxa	E. alsinoides, E. ampliata, E. arvensis, E. breviflora, E. breweri, E. caespitosa, E. cardinalis, E. decora, E. dentata, E. floribunda, E. grandis, E. guttata, E. inflatula, E. jungermannioides, E. lewisii, E. microphylla, E. moschata, E. nasuta, E. patula, E. primuloides, E. ptilota, E. pulsiferae, E. scouleri, E. suksdorfii	E. alsinoides, E. ampliata, E. arvensis, E. breviflora, E. breweri, E. caespitosa, E. decora, E. dentata, E. floribunda, E. grandis, E. guttata, E. inflatula, E. jungermannioides, E. lewisii, E. microphylla, E. moschata, E. nasuta, E. patula, E. primuloides, E. ptilota, E. pulsiferae, E. scouleri, E. suksdorfii, E. washingtonensis
Web links	Local floras: OR, WA Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA, OR, WA Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe cardinalis

Washington monkey-flower

scarlet monkey-flower

Habit

Annuals with fibrous roots or threadlike taproot; stems 5-25 cm, erect to ascending, straight or sharply bent at nodes, typically heavily branched, not angled; puberulent- to villous-glandular, flattened gland-tipped hairs 0.1-0.8 mm, sometimes transparent.

Rhizomatous perennials; stems 25-75 cm, usually erect or curving upward, branched, somewhat glandular-villous or becoming glabrous.

Leaves

Leaves cauline, basal ones not persistent;

petioles 2-14 mm;

blade triangular to ovate to narrowly ovate, 4-16 mm long and 2-11 mm broad, palmate venation, base rounded to cuneate to truncate, margins finely toothed or entire, apex acute, surfaces hairy-glandular as stems.

Leaves typically cauline; sessile;

blade ovate to elliptic to obovate, 20-90 mm long and 10-38 mm broad, palmate venation, base acuminate and nearly clasping, margins irregularly toothed, apex acute, surfaces hairy as stems.

Flowers

Axillary flowers 1-6, emerging from nodes throughout; fruiting pedicels 20-50 mm, densely covered with minute stalked glands;

calyx greenish, ridged, tubular, slightly inflated, 6-8 mm, margins toothed or lobed, covered with minute stalked glands, lobes pronounced, erect;

corollas yellow with small reddish brown dots, lower limb with two whitish patches, symmetric bilaterally, bilabiate;

tube-throat funnel-shaped and 8-10 mm, protruding beyond calyx margin;

limb expanded 7-10 mm, lobes obovate-oblong, apex rounded to somewhat cuneate;

styles hispid-hirtellous;

anthers not protruding, glabrous.

Axillary flowers 2-12, emerging from leaf axes at mid- to ends of stem; fruiting pedicels generally 30-90 mm;

calyx cylindric to nearly bell-shaped, not inflated, 17-28 mm, hairy or bristly-hairy, lobes 4-7 mm, ovate to nearly deltate, apex attenuate-acute;

corollas red to orangish red or rarely yellow, throat yellowish with red stripes, palate red, yellow-villous and not spotted or striped, symmetric bilaterally, distinctly bilabiate;

tube-throat funnel-shaped, generally 20-30 mm, protruding beyond calyx margin;

throat open;

styles glabrous;

anthers protruding, white-hairy, thecae spreading.

Fruits

Capsules 5-8.5 mm, included.

Capsules 10-16 mm, included.