Burke: Erythranthe tilingii vs. Erythranthe washingtonensis

	Erythranthe tilingii	Erythranthe washingtonensis
		Washington monkey-flower
Habit	Mat-forming perennial from well-developed, creeping rhizomes, often with stolons as well, the stems 0.5-2 dm. tall, mostly glabrous.	Annuals with fibrous roots or threadlike taproot; stems 5-25 cm, erect to ascending, straight or sharply bent at nodes, typically heavily branched, not angled; puberulent- to villous-glandular, flattened gland-tipped hairs 0.1-0.8 mm, sometimes transparent.
Leaves	Leaves opposite, mostly sessile, the blade under 2.5 cm. long, elliptic to ovate, slightly reduced upward, with a few irregular teeth, sub-palmately veined.	Leaves cauline, basal ones not persistent; petioles 2-14 mm; blade triangular to ovate to narrowly ovate, 4-16 mm long and 2-11 mm broad, palmate venation, base rounded to cuneate to truncate, margins finely toothed or entire, apex acute, surfaces hairy-glandular as stems.
Flowers	Flowers few, solitary in the leaf axils, on long pedicels; calyx 5-toothed, irregular, the upper tooth much the largest, the 2 lower ones tending to fold upward; corolla large for the size of the plant, 2-4 cm. long, strongly bilabiate, with flaring throat, yellow with maroon dots or splotches on the pubescent lower lip; stamens 4.	Axillary flowers 1-6, emerging from nodes throughout; fruiting pedicels 20-50 mm, densely covered with minute stalked glands; calyx greenish, ridged, tubular, slightly inflated, 6-8 mm, margins toothed or lobed, covered with minute stalked glands, lobes pronounced, erect; corollas yellow with small reddish brown dots, lower limb with two whitish patches, symmetric bilaterally, bilabiate; tube-throat funnel-shaped and 8-10 mm, protruding beyond calyx margin; limb expanded 7-10 mm, lobes obovate-oblong, apex rounded to somewhat cuneate; styles hispid-hirtellous; anthers not protruding, glabrous.
Fruit(s)	Capsule.	Capsules 5-8.5 mm, included.

	Erythranthe tilingii	Erythranthe washingtonensis
Flowering time	July-September	April-June
Habitat	Wet meadows and wet, rocky slopes at high elevations in the mountains.	Shallow or gravelly soils of vernally moist areas at low elevations.
Distribution	In the Cascade and Olympic Mountains of Washington; Alaska south to California, east to Montana, Colorado and New Mexico. [WildflowerSearch map]	Occurring east of the cascades crest in the Columbia River Gorge in Washington; south-central Washington to adjacent north-central Oregon. [WildflowerSearch map]
Origin		Native
Conservation status	Not of concern	Historical in Washington (WANHP)
Sibling taxa	E. alsinoides, E. ampliata, E. arvensis, E. breviflora, E. breweri, E. caespitosa, E. cardinalis, E. decora, E. dentata, E. floribunda, E. grandis, E. guttata, E. inflatula, E. jungermannioides, E. lewisii, E. microphylla, E. moschata, E. nasuta, E. patula, E. primuloides, E. ptilota, E. pulsiferae, E. scouleri, E. suksdorfii, E. washingtonensis	E. alsinoides, E. ampliata, E. arvensis, E. breviflora, E. breweri, E. caespitosa, E. cardinalis, E. decora, E. dentata, E. floribunda, E. grandis, E. guttata, E. inflatula, E. jungermannioides, E. lewisii, E. microphylla, E. moschata, E. nasuta, E. patula, E. primuloides, E. ptilota, E. pulsiferae, E. scouleri, E. suksdorfii
Web links	Local floras: BC, CA, OR Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: OR, WA Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe washingtonensis

Washington monkey-flower

Habit

Mat-forming perennial from well-developed, creeping rhizomes, often with stolons as well, the stems 0.5-2 dm. tall, mostly glabrous.

Annuals with fibrous roots or threadlike taproot; stems 5-25 cm, erect to ascending, straight or sharply bent at nodes, typically heavily branched, not angled; puberulent- to villous-glandular, flattened gland-tipped hairs 0.1-0.8 mm, sometimes transparent.

Leaves

Leaves opposite, mostly sessile, the blade under 2.5 cm. long, elliptic to ovate, slightly reduced upward, with a few irregular teeth, sub-palmately veined.

Leaves cauline, basal ones not persistent;

petioles 2-14 mm;

blade triangular to ovate to narrowly ovate, 4-16 mm long and 2-11 mm broad, palmate venation, base rounded to cuneate to truncate, margins finely toothed or entire, apex acute, surfaces hairy-glandular as stems.

Flowers

Flowers few, solitary in the leaf axils, on long pedicels;

calyx 5-toothed, irregular, the upper tooth much the largest, the 2 lower ones tending to fold upward;

corolla large for the size of the plant, 2-4 cm. long, strongly bilabiate, with flaring throat, yellow with maroon dots or splotches on the pubescent lower lip;

stamens 4.

Axillary flowers 1-6, emerging from nodes throughout; fruiting pedicels 20-50 mm, densely covered with minute stalked glands;

calyx greenish, ridged, tubular, slightly inflated, 6-8 mm, margins toothed or lobed, covered with minute stalked glands, lobes pronounced, erect;

corollas yellow with small reddish brown dots, lower limb with two whitish patches, symmetric bilaterally, bilabiate;

tube-throat funnel-shaped and 8-10 mm, protruding beyond calyx margin;

limb expanded 7-10 mm, lobes obovate-oblong, apex rounded to somewhat cuneate;

styles hispid-hirtellous;

anthers not protruding, glabrous.

Fruit(s)

Capsule.

Capsules 5-8.5 mm, included.