Burke: Erythranthe tilingii vs. Erythranthe pulsiferae

	Erythranthe tilingii	Erythranthe pulsiferae
		candelabrum monkey-flower, Pulsifer's monkey-flower
Habit	Mat-forming perennial from well-developed, creeping rhizomes, often with stolons as well, the stems 0.5-2 dm. tall, mostly glabrous.	Annuals with shallow, fibrous roots; stems 5-12 cm, erect, not bent or curved at nodes, not branched or only slightly branched proximally, covered with minute stalked glands, gland-tipped hairs 0.1-0.3.
Leaves	Leaves opposite, mostly sessile, the blade under 2.5 cm. long, elliptic to ovate, slightly reduced upward, with a few irregular teeth, sub-palmately veined.	Leaves basal and cauline; petioles 2-9 mm with 3 distinct veins and 2 wings; blade elliptic-oblong to ovate or oblanceolate, 3-14 mm long and 2-9 mm broad, palmate venation with 3 veins, base cuneate to attenuate, margins finely toothed to entire, apex acute to obtuse, surfaces glandular-hairy as stems.
Flowers	Flowers few, solitary in the leaf axils, on long pedicels; calyx 5-toothed, irregular, the upper tooth much the largest, the 2 lower ones tending to fold upward; corolla large for the size of the plant, 2-4 cm. long, strongly bilabiate, with flaring throat, yellow with maroon dots or splotches on the pubescent lower lip; stamens 4.	Axillary flowers 1-9, emerging from mid-stem to distal nodes; fruiting pedicels horizontally spreading-curved, 12-38 mm, glandular-hairy as stems; calyx cylindric, somewhat inflated, 7-10 mm, margins toothed or lobed, glandular-hairy as stems, lobes pronounced, erect; corollas yellow, tube-throat, palate ridges, lower limb yellow to light yellow, lower limb with red dots or not, symmetric bilaterally, barely bilabiate; tube-throat funnel-shaped and 6-9 mm, protruding beyond calyx margin; lobes widely obovate to somewhat orbicular, apex rounded; styles glabrous; anthers not protruding, glabrous.
Fruit(s)	Capsule.	Capsules 5-8 mm, included.

	Erythranthe tilingii	Erythranthe pulsiferae
Flowering time	July-September	May-July
Habitat	Wet meadows and wet, rocky slopes at high elevations in the mountains.	Vernally moist areas, seeps, and stream banks from the lowlands to middle elevations.
Distribution	In the Cascade and Olympic Mountains of Washington; Alaska south to California, east to Montana, Colorado and New Mexico. [WildflowerSearch map]	Occurring east of the Cascades crest in south-central Washington; south-central Washington to California, [WildflowerSearch map]
Origin		Native
Conservation status	Not of concern	Sensitive in Washington (WANHP)
Sibling taxa	E. alsinoides, E. ampliata, E. arvensis, E. breviflora, E. breweri, E. caespitosa, E. cardinalis, E. decora, E. dentata, E. floribunda, E. grandis, E. guttata, E. inflatula, E. jungermannioides, E. lewisii, E. microphylla, E. moschata, E. nasuta, E. patula, E. primuloides, E. ptilota, E. pulsiferae, E. scouleri, E. suksdorfii, E. washingtonensis	E. alsinoides, E. ampliata, E. arvensis, E. breviflora, E. breweri, E. caespitosa, E. cardinalis, E. decora, E. dentata, E. floribunda, E. grandis, E. guttata, E. inflatula, E. jungermannioides, E. lewisii, E. microphylla, E. moschata, E. nasuta, E. patula, E. primuloides, E. ptilota, E. scouleri, E. suksdorfii, E. washingtonensis
Web links	Local floras: BC, CA, OR Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA, OR, WA Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe pulsiferae

candelabrum monkey-flower, Pulsifer's monkey-flower

Habit

Mat-forming perennial from well-developed, creeping rhizomes, often with stolons as well, the stems 0.5-2 dm. tall, mostly glabrous.

Annuals with shallow, fibrous roots; stems 5-12 cm, erect, not bent or curved at nodes, not branched or only slightly branched proximally, covered with minute stalked glands, gland-tipped hairs 0.1-0.3.

Leaves

Leaves opposite, mostly sessile, the blade under 2.5 cm. long, elliptic to ovate, slightly reduced upward, with a few irregular teeth, sub-palmately veined.

Leaves basal and cauline;

petioles 2-9 mm with 3 distinct veins and 2 wings;

blade elliptic-oblong to ovate or oblanceolate, 3-14 mm long and 2-9 mm broad, palmate venation with 3 veins, base cuneate to attenuate, margins finely toothed to entire, apex acute to obtuse, surfaces glandular-hairy as stems.

Flowers

Flowers few, solitary in the leaf axils, on long pedicels;

calyx 5-toothed, irregular, the upper tooth much the largest, the 2 lower ones tending to fold upward;

corolla large for the size of the plant, 2-4 cm. long, strongly bilabiate, with flaring throat, yellow with maroon dots or splotches on the pubescent lower lip;

stamens 4.

Axillary flowers 1-9, emerging from mid-stem to distal nodes; fruiting pedicels horizontally spreading-curved, 12-38 mm, glandular-hairy as stems;

calyx cylindric, somewhat inflated, 7-10 mm, margins toothed or lobed, glandular-hairy as stems, lobes pronounced, erect;

corollas yellow, tube-throat, palate ridges, lower limb yellow to light yellow, lower limb with red dots or not, symmetric bilaterally, barely bilabiate;

tube-throat funnel-shaped and 6-9 mm, protruding beyond calyx margin;

lobes widely obovate to somewhat orbicular, apex rounded;

styles glabrous;

anthers not protruding, glabrous.

Fruit(s)

Capsule.

Capsules 5-8 mm, included.