Burke: Erythranthe tilingii vs. Erythranthe grandis

	Erythranthe tilingii	Erythranthe grandis
		large monkey-flower
Habit	Mat-forming perennial from well-developed, creeping rhizomes, often with stolons as well, the stems 0.5-2 dm. tall, mostly glabrous.	Rhizomatous perennials, occasionally rooting at nodes nearest base; stems usually 50-120 cm, erect or decumbent basally, branched, commonly hollow, densely hirsute to soft-hirtellous to long-hairy, hairs often crinkly, glandular or not, or sometimes villous-glandular without hirtellous hairs.
Leaves	Leaves opposite, mostly sessile, the blade under 2.5 cm. long, elliptic to ovate, slightly reduced upward, with a few irregular teeth, sub-palmately veined.	Leaves basal and cauline, basal leaves generally not persistent; petioles 10-80 mm, becoming reduced distally; blade ovate to broadly elliptic, 25-60 mm long and 20-40 cm broad, palmate or nearly pinnate venation with 5-7 veins, base truncate to nearly cruneate to nearly cordate, margins crenulate to toothed, occasionally sublyrate near base, apex rounded to obtuse, surfaces of distalmost leaves hairy as stems.
Flowers	Flowers few, solitary in the leaf axils, on long pedicels; calyx 5-toothed, irregular, the upper tooth much the largest, the 2 lower ones tending to fold upward; corolla large for the size of the plant, 2-4 cm. long, strongly bilabiate, with flaring throat, yellow with maroon dots or splotches on the pubescent lower lip; stamens 4.	Inflorescence racemose, bracteate, flowers 8-26; fruiting pedicels 10-35 mm, hairy as stems; calyx straight-erect or nodding at 45-100 degrees, ovate to bell-shaped, inflated, compressed along sagittal plane, 15-22 mm, hairy as stems, throat closing; corollas yellow with red spots inside, symmetric bilaterally, bilabiate; tube-throat widely funnel-shaped, 16-24 mm, protruding 10-15 mm beyond calyx margin, limb widely expanded; styles hirtellous; anthers not protruding, glabrous.
Fruit(s)	Capsule.	Capsules 8-12 mm, included.

	Erythranthe tilingii	Erythranthe grandis
Flowering time	July-September	May-August
Habitat	Wet meadows and wet, rocky slopes at high elevations in the mountains.	Coastal areas including bluffs, dunes, wet cliff faces, marshes, ditches, lake and pond margins, and stream banks.
Distribution	In the Cascade and Olympic Mountains of Washington; Alaska south to California, east to Montana, Colorado and New Mexico. [WildflowerSearch map]	Occurring west of the Cascades crest in Washington; Washington to California. [WildflowerSearch map]
Origin		Native
Conservation status	Not of concern	Not of concern
Sibling taxa	E. alsinoides, E. ampliata, E. arvensis, E. breviflora, E. breweri, E. caespitosa, E. cardinalis, E. decora, E. dentata, E. floribunda, E. grandis, E. guttata, E. inflatula, E. jungermannioides, E. lewisii, E. microphylla, E. moschata, E. nasuta, E. patula, E. primuloides, E. ptilota, E. pulsiferae, E. scouleri, E. suksdorfii, E. washingtonensis	E. alsinoides, E. ampliata, E. arvensis, E. breviflora, E. breweri, E. caespitosa, E. cardinalis, E. decora, E. dentata, E. floribunda, E. guttata, E. inflatula, E. jungermannioides, E. lewisii, E. microphylla, E. moschata, E. nasuta, E. patula, E. primuloides, E. ptilota, E. pulsiferae, E. scouleri, E. suksdorfii, E. washingtonensis
Web links	Local floras: BC, CA, OR Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: CA, OR, WA Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Erythranthe grandis

large monkey-flower

Habit

Mat-forming perennial from well-developed, creeping rhizomes, often with stolons as well, the stems 0.5-2 dm. tall, mostly glabrous.

Rhizomatous perennials, occasionally rooting at nodes nearest base; stems usually 50-120 cm, erect or decumbent basally, branched, commonly hollow, densely hirsute to soft-hirtellous to long-hairy, hairs often crinkly, glandular or not, or sometimes villous-glandular without hirtellous hairs.

Leaves

Leaves opposite, mostly sessile, the blade under 2.5 cm. long, elliptic to ovate, slightly reduced upward, with a few irregular teeth, sub-palmately veined.

Leaves basal and cauline, basal leaves generally not persistent;

petioles 10-80 mm, becoming reduced distally;

blade ovate to broadly elliptic, 25-60 mm long and 20-40 cm broad, palmate or nearly pinnate venation with 5-7 veins, base truncate to nearly cruneate to nearly cordate, margins crenulate to toothed, occasionally sublyrate near base, apex rounded to obtuse, surfaces of distalmost leaves hairy as stems.

Flowers

Flowers few, solitary in the leaf axils, on long pedicels;

calyx 5-toothed, irregular, the upper tooth much the largest, the 2 lower ones tending to fold upward;

corolla large for the size of the plant, 2-4 cm. long, strongly bilabiate, with flaring throat, yellow with maroon dots or splotches on the pubescent lower lip;

stamens 4.

Inflorescence racemose, bracteate, flowers 8-26; fruiting pedicels 10-35 mm, hairy as stems;

calyx straight-erect or nodding at 45-100 degrees, ovate to bell-shaped, inflated, compressed along sagittal plane, 15-22 mm, hairy as stems, throat closing;

corollas yellow with red spots inside, symmetric bilaterally, bilabiate;

tube-throat widely funnel-shaped, 16-24 mm, protruding 10-15 mm beyond calyx margin, limb widely expanded;

styles hirtellous;

anthers not protruding, glabrous.

Fruit(s)

Capsule.

Capsules 8-12 mm, included.